, in the headwaters of the Mokelumne River The Mokelumne River
watershed drains in parts of
Alpine,
Amador,
Calaveras,
San Joaquin, and
Sacramento Counties. Elevations range from sea level at the confluence with the San Joaquin River to at the crest of the Sierra Nevada in the headwaters of the North Fork. The Mokelumne watershed divide borders the basins of the
American River on the north, the
Calaveras and
Stanislaus rivers to the south, and the
Carson River to the east. The American, Calaveras and Stanislaus rivers, like the Mokelumne, drain westwards into the Central Valley, while the Carson flows eastwards into the
Great Basin. Drainage within the Mokelumne watershed generally occurs from east to west with all of the
perennial streams originating in the Sierra Nevada. The Mokelumne River and its headwater forks above Lodi drain the southernmost part of the basin. The part of the basin above Pardee Dam is referred to as the Upper Mokelumne River Basin and drains about , or 25.7% of the total watershed. Dry Creek, which itself is formed by four Sierra streams – Jackson, Sutter, Amador and Rancheria Creeks – drains most of the area between the upper Mokelumne and Cosumnes basins, or about . The upper watershed is situated principally in Alpine, Amador and Calaveras Counties and consists mostly of wilderness managed under various federal designations. Most of the Mokelumne and Cosumnes River headwaters lie within the Stanislaus and Eldorado National Forests. Much of the upper watershed is also protected under the
Mokelumne Wilderness. The Salt Springs State Game Refuge is located along the Mokelumne River near Salt Springs Reservoir. There are also significant private holdings in the upper basin including commercial timber land and protected watershed areas administered by the
East Bay Municipal Utility District, which diverts urban and industrial water from Pardee Reservoir. The lower watershed, mainly within San Joaquin and Sacramento Counties, is part of the rich agricultural region of the Central Valley. Grapes are the major crop grown in the Mokelumne River watershed; as of 2001,
vineyards comprised 51% of the basin's farmland. Of the remaining land, 31% is used for
livestock, 9% for orchards, and 8% for annual crops. About in the lower basin are part of the
Mokelumne River AVA. In May 2021, it was reported that a trove of 5- to 10-million-year-old fossils was unearthed in June 2020 in the Mokelumne River watershed near
Valley Springs, California, including a two-tusked
mastodon, a four-tusked
gomphothere,
rhinoceros,
camel,
horse,
tortoise,
tapir, bird, fish, and other specimens. ==History==