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Simplicidentata

Simplicidentata is a group of mammals that includes the rodents and their closest extinct relatives. The term has historically been used as an alternative to Rodentia, contrasting the rodents with their close relatives the lagomorphs. However, Simplicidentata is now defined as including all members of Glires that share a more recent common ancestor with living rodents than with living lagomorphs. Thus, Simplicidentata is a total group that is more inclusive than Rodentia, a crown group that includes all living rodents, their last common ancestor, and all its descendants. Under this definition, the loss of the second pair of upper incisors is a synapomorphic feature of Simplicidentata. The loss of the second upper premolar (P2) has also been considered as synapomorphic for Simplicidentata, but the primitive simplicidentate Sinomylus does have a P2.

Literature cited
• Landry, S.O., Jr. 1999. A proposal for a new classification and nomenclature for the Glires (Lagomorpha and Rodentia). Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reihe 75(2):283–316. • McKenna, M.C. and Bell, S.K. 1997. Classification of Mammals: Above the species level. New York: Columbia University Press, 631 pp. • McKenna, M.C. and Meng, J. 2001. A primitive relative of rodents from the Chinese Paleocene. Journal of Vertebrate Paleontology 21(3):565–572. • Meng, J. and Wyss, A.R. 1994. Enamel microstructure of Tribosphenomys (Mammalia, Glires): Character analysis and systematic implications. Journal of Mammalian Evolution 2(3):185–203. • Meng, J. and Wyss, A.R. 2001. The morphology of Tribosphenomys (Rodentiaformes, Mammalia): phylogenetic implications for basal Glires. Journal of Mammalian Evolution 8(1):1–71. • Meng, J., Hu, Y. and Li, C. 2003. The osteology of Rhombomylus (Mammalia, Glires): implications for phylogeny and evolution of Glires. Bulletin of the American Museum of Natural History, 275:1–247. • Meng, J., Kraatz, B.P., Wang, Y., Ni, X., Gebo, D.L. and Beard, K.C. 2009. A new species of Gomphos (Glires, Mammalia) from the Eocene of the Erlian Basin, Nei Mongol, China. American Museum Novitates 3670:1–11. • Rose, K.D. 2006. The Beginning of the Age of Mammals. Baltimore: The Johns Hopkins University Press, 428 pp. • Szalay, F.S. 1999. [Review of Classification of Mammals: Above the Species Level] (subscription required). Journal of Vertebrate Paleontology 19(1):191–195.
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