The biota of the formation comprise mostly marine microinvertebrates and algae, with the presence of
ammonites and abundant pollen in the Middle Toarcian sections.
Nanoplankton is one of the most representative finds of the layers, appearing deposited with interruptions, probably due to changes on the oxygen content of the sea floor.
Phytoplankton includes abundant
dinoflagellate cysts and
acritarchs, specially the genus
Micrhystridium. There registered a transgression on the sea level on the Lower Toarcian, with a rise, giving the Umbria-Marche Basin pelagic conditions, and a
regression during the Middle Toarcian. Those changes implicate the disappearance of genera of dinoflagellates from the strata to be replaced with new ones. Some dinoflagellate genera such as
Mendicodinium are more abundant, being the dominant palynological residue recovered in some samples. Foraminiferal analysis had been done, revealing changes in the salinity, with
Spirillina as the most abundant genus found, followed by
Prodentalina,
Eoguttulina,
Lenticulina,
Nodosaria,
Lingulonodosaria and
Pseudonodosaria. Invertebrate remains are mainly
brachiopods (
Pseudokingena,
Nannirhynchia and
Lokutella, among others) and
ammonites;
Eodactylites,
Pleuroceras,
Canavaria,
Trinacrioceras,
Lioceratoides, and
Praepolyplectus as predominant genera. Other marine fauna includes
Holothuriidan
Sclerites, unidentified
crinoids, fragmentary
asteroideans and fish teeth & complete specimens, including the genus
Leptolepis. There are terrestrial fossils and sediments, specially clay. The main source area for the clay could have been the palaeosoils developed on the Carbonate
Latium-Abruzzo Platform, although more recent studies prove a continental origin. The Main fossils from terrestrial deposits are plant Pollen, mostly
Pteridophyta spores and
Circumpolles, and
Palynomacerals. The genera includes
Leptolepidites (
Lycopsida) spp.,
Trilites (
Filicopsida) spp.,
Ischyosporites (
Pteridopsida) spp. or
Foveosporites (
Selaginellaceae) spp. and others less abundant, such as the genus
Callialasporites (
Coniferales). There is a curious reduction of the
Cheirolepidiaceae genus
Classopollis, abundant on coeval associations from northern Europe,
Portugal, northeast
Italy and
Israel, as a possible effect of paleoclimatic changes.
Megaflora Several plant leaves and fragments of wood were not identified. == See also ==