In the wake of both Gus Portokalos and Ian's father Rodney's passing a year ago, the Portokalos family is grappling with grief. Nick, now separated, returns home, displaying immature behavior. Athena and her husband Yiannis are caring for Maria, who is battling dementia. Toula and Ian, though happily married, are facing the challenges of parenthood while their daughter Paris struggles with the responsibility of college, completely unbeknownst to them. Victory, a distant relative and the mayor of Gus' hometown Vrisi, invites the family for a supposed reunion. On the flight to Greece, Aunt Voula introduces her assistant Aristotle, Paris' college-mate, and Toula reveals she's bringing Gus' journal, containing his life story, to give to his three childhood friends in Greece as promised. They arrive in Athens, where Victory greets them, and go sightseeing. In the nearly abandoned Vrisi, they find Gus' childhood home and meet Alexandra, Gus's old flame. There, Victory confesses there's no reunion, hoping the Portokaloses would revive the village. The family goes to a nearby town the following day, where Toula searches for Gus's friends without success and gets drunk on
ouzo. Nick searches for the village's oldest tree and Ian befriends a monk, who possesses information about everyone's activities in the village. Meanwhile, Aristotle tries to communicate with Paris, who rebuffs his advances without explaining why. At dinner, Toula and Ian meet Peter, Alexandra's son by Gus, whom the latter never knew about, as well as his son Christos, and Alexandra's assistant Qamar. Afterwards, Christos and Qamar take Aristotle and Paris to a club, where they reveal they are secretly betrothed. The next day, Toula informs her family about Peter, but struggles to locate Gus' friends. Fortunately, the monk gave Ian a list of those who left the village, pinpointing one who moved to
Syros. Toula shares this with Nikki and Angelo, who head to Syros and return with them. Meanwhile, Toula discovers Nick brought Gus's ashes to Greece, following his father's wish for them to be scattered by the oldest tree in Vrisi. Later that evening, the family learns about Christos and Qamar's engagement, coupled with the revelation of Paris being on academic probation. Peter and Alexandra express disapproval, prompting Ian to offer intervention for Paris. However, the younger generation rejects external interference, opting to address their issues independently. The next day, Peter comes to terms with Christos and Qamar's relationship, with Alexandra eventually doing the same. Victory recognizes the potential of a wedding to revitalize the village and initiates plans. Toula and Ian receive encouragement to unwind and relish their vacation while the rest of the family collaborates in organizing the wedding, albeit with their assistance in the final stages. In a joyous celebration, Qamar's family joins them for her wedding to Christos. Paris opens up about her split with Aristotle, citing struggles with independence, and they agree to take things slow, much to Aunt Voula's delight. Nikki and Angelo arrive with Gus' friends, and Toula presents them with his journal. Later, Toula, Nick, and Peter privately scatter Gus' ashes around the tree, and Victory's mayorship receives praise. Toula and Nick decide to bring Peter with them to America, now as part of the family. The siblings, on the way, engage in a playful dispute over who should now lead the family, each citing their actions during the journey. ==Cast==