These groups are sometimes called "lower plants", referring to their status as the earliest plant groups to evolve, but the usage is imprecise since both groups are
polyphyletic and may be used to include
vascular cryptogams, such as the
ferns and
fern allies that reproduce using spores. Non-vascular plants are often among the first species to move into new and inhospitable territories, along with
prokaryotes and
protists, and thus function as
pioneer species. Mosses and leafy liverworts have structures called
phyllids that resemble
leaves, but only consist of single sheets of cells with no internal air spaces, no
cuticle or
stomata, and no xylem or phloem. Consequently, phyllids are unable to control the rate of water loss from their tissues and are said to be
poikilohydric. Some liverworts, such as
Marchantia, have a cuticle, and the sporophytes of mosses have both cuticles and stomata, which were important in the
evolution of land plants. All
land plants have a
life cycle with an
alternation of generations between a diploid
sporophyte and a haploid
gametophyte, but in all non-vascular land plants, the gametophyte generation is dominant. In these plants, the sporophytes grow from and are dependent on gametophytes for supply of water and mineral nutrients and photosynthate, the products of
photosynthesis. ==Environmental role==