It is now considered a member of the genus
Eurycea, but was originally described as
Typhlotriton nereus. It was described in 1968, but was later synonymized with the
grotto salamander (
E. spelaea), but a 2017 study found substantial genetic differences between the clades classified in
E. spelaea and once again split them into distinct species. It is thought to have diverged from the
southern grotto salamander (
E. braggi) during the Late
Miocene. All three grotto salamanders are thought to descend from an ancestral surface-dwelling form. == Distribution and habitat ==