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Nymphaea nouchali var. caerulea

Nymphaea nouchali var. caerulea, or simply Nymphaea caerulea, also known as blue lotus or blue water lily among many other names, is a water lily in the genus Nymphaea, a botanical variety of Nymphaea nouchali.

Description
Vegetative characteristics This is an aquatic (euhydrophyte) herb with a tuberous rhizome. That is to say, it has small tubers that may develop into short vertical rhizomes. It is a perennial. One plant can spread over an area of about 1 metre. Nymphaea caerulea contains the psychoactive alkaloid nuciferine, which contributes to its euphoric effects. Chemical analyses using mass spectrometry have shown that authentic flowers have significantly higher concentrations of nuciferine compared to other water lilies. ==Cytology==
Cytology
The chromosome count is n = 14. The genome size is 567.24 Mb. ==Taxonomy==
Taxonomy
Nymphaea spectabilis, a purple form known from cultivation, and N. capensis, found throughout eastern, central and southern Africa, as well as a number of other named taxa, were synonymised to N. nouchali var. caerulea in the 1989 addition to the Flora of Tropical East Africa (FTEA) series, a position which has generally been accepted, although some of the authorities in Bangladesh When genomes from the water lily genus (Nymphaea) were published in the journal Nature in 2020, N. caerulea was cited under that name, not as N. nouchali var. caerulea. Another phylogenetic study from 2021 found N. caerulea (as N. capensis) to be closest related to N. colorata, an east African species. Nymphaea nouchali is itself a taxonomically challenging species, with a distribution that spans Australia, throughout southern Asia, across Africa to the Western Cape. This subgenus appears to be phylogenetically sound. ==Distribution==
Distribution
The native distribution covers North Africa along the Nile and south throughout central, East and Southern Africa. Congo-Brazzaville, Angola but the plants in the wild are now thought to be N. caerulea. It is considered an environmental weed in Australia. ==Ecology==
Ecology
It has a habitat consisting of rivers, lakes and pools. The waterlily stands in this lake are especially poor in invertebrate biodiversity, which may reflect the low levels of dissolved oxygen near the sediments in this habitat. In Lake Bisina, Uganda, N. caerulea is most clearly associated with Utricularia reflexa; this may be due to similar ecological niches, it may just mean the small, rootless, free-moving Utricularia simply get snagged on the petioles, but it may indicate some sort of a commensal relationship, with U. reflexa being shaded by the leaves of N. caerulea. Hydrilla verticillata is another plant which seems to sometimes occur together with the waterlily in this lake, as well as in Lake Bunyonyi. Pollination is entomophilous. In fact, the carpellary appendages in this type of water-lily appear to have evolved specifically to attract bee species in general. In a way, these waterlilies are parasites of the services of bees, attracting the insects by deceit, without actually rewarding them for their labours. These seeds only germinate when heavy rains flood the banks, and they are submerged under a layer of water. In Kenya, N. caerulea is positively associated with the freshwater snail Biomphalaria pfeifferi, which is a main host of human schistosomiasis. The edible American crayfish Procambarus clarkii eliminates the mollusc, as well as feeding on the water-lily. The crayfish was first introduced to Kenya in 1966 as a species with which to enhance the local fisheries. In Lake Naivasha, N. caerulea was extremely common until the 1970s, and there is still a seedbank around the shores of the lake. Procambarus clarkii was introduced to the lake in 1970, and now supports an annual harvest of a few thousand kilograms, but it may have been responsible for eliminating not only the water-lily in the main lake by 1983, but all native aquatic plant species in this water body. It is not the only potential culprit; invasive mats of exotic floating vegetation have also taken over the lake, two different commercially fishable fish species have been introduced, and the new fisheries upon these three species could all be responsible, or a combination. ==Uses==
Uses
The rootstock of the blue water lily was collected and eaten in western South Africa around 1800, either raw or in curries, in particular by the Cape Malays and farming communities in the Cape, although this practice has now died out. Psychoactive effects It has been suggested that Nymphaea caerulea was used in ancient Egypt for religious rituals, sexual enhancement, and other purposes, due to the purported presence of nuciferine and apomorphine, the latter of which is also used today to treat erectile dysfunction. The key active constituents and mechanisms of action of the plant are unknown. Andrew Sherratt and colleagues investigated the effects of Nymphaea caerulea in two human volunteers in the 1998 television series Sacred Weeds. It was the first known study and demonstration of the psychoactive effects of the plant in humans. since 1963. with photos of a real Nymphaea caerulea posted on their social media as recent as 2019. Due to its colour, it was identified, in some beliefs, as having been the original container, in a similar manner to an egg, of Atum, and in similar beliefs Ra, both solar deities. As such, its properties form the origin of the "lotus variant" of the Ogdoad cosmogony. It was also the symbol of the Egyptian deity Nefertem. Often depicted in ancient art and found in tombs—including the petals discovered in King Tutankhamun's burial chamber in 1922—it held a sacred role in religious rituals. Notably featured in the Festival of Drunkenness honoring Hathor, goddess of love and fertility, the flower was believed to induce visions when soaked in wine, possibly used in ecstatic or hallucinogenic rites. Native to the Nile River, the species has become rare due to ecological changes, particularly following the construction of the Aswan Dam, and is now considered threatened. Toxicity and health effects While unregulated in the United States and sold as a dietary supplement, consumption of Nymphaea caerulea can cause toxic effects. Modern use often involves drinking infusions (water or alcohol based) made from the plant material or inhaling extracts using electronic cigarettes (vaping). With the latter being a more rapid and efficient method, delivering higher effective doses to the body and increasing the potential for toxicity. A 2021 case series described five patients who presented to an emergency department with altered mental status after consuming blue lotus. Symptoms included sedation, disorientation, and perceptual disturbances. Other reported effects were tachycardia, anxiety, and, in one case, a persistent erection (priapism). All patients in the series recovered with supportive care, such as intravenous fluids and observation, without needing sedating medications. The psychoactive symptoms typically resolved within 3 to 4 hours. Currently, standard urine drug screens do not detect blue lotus, complicating the diagnosis. ==Legal issues==
Legal issues
United States While Nymphaea caerulea is not a federally controlled substance in the United States according to the Drug Enforcement Administration (DEA) and is therefore legal to sell in most states, it is explicitly prohibited for use by all members of the United States Armed Forces. The plant is included on the Department of Defense Prohibited Dietary Supplement Ingredients List, and its use is a violation of the Uniform Code of Military Justice (UCMJ) The plant was banned in Poland in March 2009. Possession and distribution lead to a criminal charge. N. caerulea has been illegal in Russia since April 2009, along with related products such as Salvia divinorum, Argyreia nervosa, and others. ==See also==
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