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Octopus tetricus

Octopus tetricus, the gloomy octopus, the common Sydney octopus, or the peachy octopus, is a species of octopus from the subtropical waters of eastern Australia and northern New Zealand. O. tetricus belongs to the Octopus vulgaris species group. All species within the O. vulgaris group are similar in morphology, behaviour, and physiology. The English translation of O. tetricus (Latin) is 'the gloomy octopus'. It is a significant species in the fishing industry in Australia. They play an important role in energy flux between trophic levels in the marine environment.

Distribution
O. tetricus was originally discovered in New South Wales and was also found along the eastern Australian coastline. It occurs from Lakes Entrance in Victoria O. tetricus is distributed in the subtropical seas of eastern Australia and northern New Zealand, including Lord Howe Island. A close relative, Octopus djinda, occurs at similar latitudes in Western Australia, from Shark Bay to Cape Le Grand, O. tetricus populations in New Zealand are found along the North-East coast of the North Island, between the Bay of Plenty and Northland. It is hypothesised that warmer waters spreading south of Australia are influencing populations to O. tetricus to disperse further south in response to climate change. == Anatomy and morphology ==
Anatomy and morphology
Octopus tetricus is normally coloured grey to mottled brown with rufous arm faces that taper towards the tip. They have orange colouration of arms, funnel, and mantle opening. and a mantle length of up to . The brain, optic lobes, and the highly developed arm nervous system are the three main components of the octopus nervous system. Each arm of the octopus has been described as having its own brain, where individual processing, motor systems, and sensory exploration can occur. This is where the concept that octopus have many brains comes from. The arm nervous system of the octopus contains three-fifths of the octopus's total number of neurons – 350 million neurons distributed equally throughout each of its eight arms. == Taxonomy and etymology ==
Taxonomy and etymology
Octopus tetricus Gould, 1852 was described by American conchologist Augustus Addison Gould in the 1852 publication, Mollusca of the United States Exploring Expedition under Captain Charles Wilkes. An illustration of O. tetricus was also published a few years later (plate 47). The type specimen is listed as being from Sydney, New South Wales, Australia. The New Zealand species O. gibbsi is considered to be a synonym of O. tetricus. The species epithet tetricus means 'forbidding' or 'gloomy'. == Life cycle ==
Life cycle
Reproduction and growth The reproductive cycle of females is tied to seasonal changes, similar to many other species. Females reach maturity around Australia's spring and summer seasons in order to mate and lay eggs. During the mating process the male O. tetricus passes spermatophores to the female in two different ways. The male either wraps his arms and web around the female's mantle or reaches his mating arm from a distance and inserts it into the female's mantle. The males third right arm is the specialised mating arm, and the sperm is moved down a groove on this arm which has a sucker-less tip. O. tetricus start out as eggs that are laid in large numbers in the octopus's nest, approximately 270,000 eggs per female. Female O. tetricus have also been known to store viable spermatozoa for up to 114 days. The amount of time embryonic development takes varies with water temperature, and newly hatched O. tetricus larvae are about 2.5 mm long and 1.1 mm wide, approximately the size of a grain of rice. These larvae go through a stage called the paralarval stage where they are considered planktonic, or free floating, organisms before they settle to the bottom and grow large enough to hunt for their food. This stage may last around 35–60 days. The females of O. tetricus have been known to cannibalise the males following mating. Temperature plays a key role in growth of this octopus species. With a good food supply, octopuses that reside in areas with a cooler water temperature tend to grow slower during the key growth phase and when they reach maturity they are generally larger than octopuses that are found in warmer water temperatures. Lifespan Adult O. tetricus is observed to have a relatively small body size and a lifespan of approximately 11 months, although there is evidence that they can live up to 2 years. They have a very fast growth rate, reaching adult size after approximately 100 days. Female O. tetricus rarely eat or sleep during the protection of the nest, devoting all resources to nourishing its offspring, and die shortly after the eggs hatch. Females are found to mature at a slower rate and become larger than male O. tetricus. This is a semelparous species, meaning each female reproduces once in her lifetime. == Habitat and behavioural ecology ==
Habitat and behavioural ecology
Habitat Octopus tetricus occurs in the intertidal zone along rocky shores and seagrass beds, and in the ocean, out up to , O. tetricus has been observed shifting and transporting sediment and shells to maintain their dens. They tend to have a preference for materials that are free from barnacles and encrustations, although the scale on which this is the case is dependent on foraging success. A solid object can serve as a good den that can also be the start of a new settlement for O. tetricus. Studies show that O. tetricus has higher populations in patch reef habitats than broken reef habitats, and were scarcely found on flat reefs. Adult O. tetricus were also found to occupy coastal reefs in the summer and then disappear around the second week of April, which is the second week of autumn in Australia. Shelters serve a vital role in octopus ecology. When resources are scarce, such as food and dens, they will often attempt to exclude and displace other individuals. It is a territorial species which sits out the day in a lair among rocks and rubble, the rubble being collected to create a defensible lair. The lairs of this species can be identified by the shells of the octopus's prey which it scatters around its home. They move about the rocks by crawling using their arms but they can use their siphon to propel themselves through the water by generating a jet of water or to move (throw) shells, silt, and algae, sometimes targeting other individual octopus. Throwing is a very rare and uncommon behaviour exhibited in non-humans, only seen in a few other species such as non-human primates and dolphins. Females have also been observed storing sperm for up to 3 months after mating, before laying any eggs. The species is generally known to be solitary, but complex social behaviours have been observed by scientists. Animals that predate on O. tetricus include some species of dolphin, the short-tail stingray (Dasyatis brevicaudata), southern eagle rays (Myliobatis goodei), wobbegong sharks, and some fish. Fish, however, usually attack when in schools, such as Chinaman leather-jackets. There is no evidence to suggest that any one species solely relies on O. tetricus as a food source. Generally, O. tetricus are considered a solitary, crepuscular species, meaning they emerge from their dens to hunt and feed alone at dusk and dawn before returning to the safety of their enclosures to consume prey and to spend their daytime. Contrary to this, it is believed that the solitary, nocturnal behaviour is context-dependent. In high densities of the species, individuals become much more social and active during the day. This could be due to a greater sense of safety and protection when in larger groups. Differences in behaviour between sexes of the species are apparent, such as males being less likely to modify dens and shelters compared to females. Using properties of skin is a significant element to both combative and courtship types of behaviour. They are also able to communicate with other octopus without disrupting their crypsis, meaning they do not have to be vulnerable in order to communicate with each other. Changes in skin properties can be used to signal that they feel threatened, for example when faced with a predator, or when a female rejects a males advances for mating. However, it does not play a role in females selecting male mating partners. == Fisheries ==
Fisheries
Octopus tetricus may be caught as bycatch in trawl and lobster-pot fisheries and is then sold for both human consumption and for use as bait. The species made up to 43% of total cephalopod fishing catches in 2007, however, it is possible that O. tetricus was sometimes misidentified as Octopus australis. Since 2016, annual catches of the species has increased over 50% to 12.6 tonnes. In 2017, it was said that O. tetricus made up 14% of the total octopus catch. Their catch rates are highest during spring and summer, by which they then drop significantly just as autumn arrives. Historically and through to the present day, the species is a significantly important species in the fishing industry across Australia. == Conservation status ==
Conservation status
Octopus tetricus is listed as 'Least Concern' on The IUCN Red List of Threatened Species. == References ==
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