During the
Maastrichtian the inner area was situated within an anoxic marine setting, as revelated by bituminous, very oily and dolomitic black shells. Palynology of the underliying and coeval
El Koubbat Formation has revelated an microbiota composed of
Dinoflagellates,
Pterospermopsis,
Scolecodonts and
Tasmanaceae. The formation reflects a shallow marine to lagoonal setting, transitioning from fine sands to marls and limestones. The presence of marine organisms, including bivalves and gastropods, along with phosphatic layers containing fish remains, indicates periodic marine incursions, with intermittent lagoonal or nearshore conditions. Oxidation and sedimentary discontinuities suggest episodes of exposure or subaerial erosion. The overall environment was likely a shallow, low-energy, marginal marine setting with occasional deeper marine influences. Marine Amniote fauna is known from coeval layers of the
El Koubbat Formation, mostly
Mosasaurs (
Halisaurinae,
Gavialimimus,
Globidens,
Mosasaurus,
Prognathodontini) and
Elasmosauridae plesiosaurs. == Biota ==