Ophthalmosaurus

of O. icenicus Ophthalmosaurus was a medium-sized ichthyosaur, growing to measure in length and weighing between . == Discovery and species ==

Ophthalmosaurus was first described by Harry Seeley in 1874 with particular focus on the morphology of the clavicular bones. Over the years following its description a variety of genera have been sunk into Ophthalmosaurus. Among them, Apatodontosaurus, Ancanamunia, Baptanodon, Mollesaurus, Paraophthalmosaurus, Undorosaurus and Yasykovia were all considered junior synonyms of Ophthalmosaurus in a study published by Maisch & Matzke in 2000. However, more recent cladistic analyses have contested Maisch & Matzke's conclusion. Mollesaurus periallus from Argentina was considered a valid genus of ophthalmosaurid by Druckenmiller and Maxwell (2010), Paraophthalmosaurus and Yasykovia were both recovered as distinct genera by Storrs et al., but were later sunk into Nannopterygius while Undorosaurus's validity is now accepted by most authors, including Maisch (2010) who originally proposed the synonymy. The two other Russian taxa might be also valid. Ophthalmosaurus natans was described as Sauranodon, then later renamed to Baptanodon by Marsh in 1880. However this decision was questioned not long afterwards with Baptanodon instead being considered an American species of Ophthalmosaurus. Recent analysis have recovered the species as closer to other ophthalmosaurines than to the Ophthalmosaurus type species, suggesting that the previous name should be reinstated. Similarly, Ophthalmosaurus chrisorum, whose holotype has been recovered in Canada and described by Russell in 1993, was moved to its own genus Arthropterygius in 2010 by Maxwell. While primarily known from the Jurassic, material from the Spilsby Sandstone dating to the early Berriasian stage of the Lower Cretaceous has been referred to cf. Ophthalmosaurus (i.e., either Ophthalmosaurus or a closely related species). == Classification ==

Within Ophthalmosauridae, Ophthalmosaurus was once considered most closely related to Aegirosaurus. However, many recent cladistic analyses found Ophthalmosaurus to nest in a clade with Acamptonectes and Mollesaurus. Aegirosaurus was found more closely related to Platypterygius, and thus does not belong to the Ophthalmosaurinae. Phylogeny The cladogram below follows Fischer et al. 2012. }} The following cladogram shows a possible phylogenetic position of Ophthalmosaurus in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020). }} == Palaeobiology ==

Ophthalmosaurus icenicus possessed small teeth with robust tooth crowns and signs of slight wear differing notably from the robust teeth of later species of Platypterygius, known to have hunted large prey including turtles and birds, and the minute teeth of Baptanodon, interpreted to be a soft prey specialist. Fischer et al. (2016) conclude that this intermediary tooth morphology indicates that Ophthalmosaurus icenicus was most likely a generalist predator, feeding on a variety of smaller prey items. Ophthalmosaurus could likely dive for around 20 minutes. Assuming a conservative cruising speed of ( being more likely), Ophthalmosaurus could reach depths of or more during a dive, reaching the mesopelagic zone. However, while studies on the biomechanics of Ophthalmosaurus suggests that such feats could be physically achieved, studies on the environment of the Peterborough member of the Oxford Clay suggest that Ophthalmosaurus instead inhabited relatively shallow waters there, being determined to have been just deep at a distance of from the shore. == See also ==