Oryza sativa contains two major subspecies: the sticky, short-grained
japonica or sinica variety, and the nonsticky, long-grained ''
rice variety. Japonica
was domesticated in the Yangtze Valley 6,000–9,000 years ago, and its varieties can be cultivated in dry fields (it is cultivated mainly submerged in Japan), in temperate East Asia, upland areas of Southeast Asia, and high elevations in South Asia, while indica'' was domesticated around the
Ganges 4,500–8,500 years ago, A third subspecies, which is broad-grained and thrives under tropical conditions, was identified based on morphology and initially called
javanica, but is now known as
tropical japonica. Examples of this variety include the medium-grain 'Tinawon' and 'Unoy' cultivars, which are grown in the high-elevation rice
terraces of the
Central Cordillera Mountains of northern
Luzon, Philippines. Glaszmann (1987) used
isozymes to sort
O. sativa into six groups:
japonica,
aromatic,
indica,
aus,
rayada, and
ashina. Garris
et al. (2004) used
simple sequence repeats to sort
O. sativa into five groups:
temperate japonica,
tropical japonica and
aromatic comprise the
japonica varieties, while
indica and
aus comprise the
indica varieties. The Garris scheme has held up against newer analyses as of 2019, though one 2014 article argues that
rayada is distinct enough to be its own group under
japonica.
Etymology The generic name
Oryza is a classical Latin word for rice, while the specific epithet
sativa means "cultivated". == Genetics ==