Marsh grass shrimp have been investigated for their maneuverability and swimming. These shrimp swim
metachronally, creating an appendage wave, starting with the
pleopod closest to the tail, beating all of their appendages with a phase lag. They also modulate the profile area of their appendages to create net thrust. Through
Particle Image Velocimetry, two drag-reducing mechanisms have been identified in marsh shrimp swimming that enable them to reduce drag during the return stroke of their beat. The first of those mechanisms is asymmetric flexibility, with the appendages staying mostly rigid during the power stroke and being flexible during the return stroke, this results in a reduced wake. The second of those mechanisms is grouping of the appendages. ==References==