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Blackbelly lanternshark

The blackbelly lanternshark or lucifer shark is a shark of the family Etmopteridae found around the world in tropical and temperate seas at depths between 150 and 1,250 meters – the mesopelagic zone. Compared to other mesopelagic fish predators and invertebrates, the blackbelly lanternshark is thought to reside in shallower, more southern waters. E. lucifer can reach up to 47 centimeters in length and consumes mesopelagic cephalopods, fish, and crustaceans. Blackbelly lanternsharks are bioluminescent, using hormone controlled mechanisms to emit light through ventral photogenic organs called photophores and are presumed to be ovoviviparous. The blackbelly lanternshark has been classified as "Not Threatened" within the New Zealand Threat Classification System.

Distribution and habitat
Etmopterus lucifer is prevalent in New Zealand's deep waters, most commonly found on the south Chatham Rise, with the greatest occurrence recorded at around 500 meters in depth. E. lucifer is categorized as a mesopelagic fish, preferring shallower and more southern habitats. Between 1992 and 2010, a significant increase in E. lucifer biomass was observed along the east coast of the North Island. In New Zealand's deep-sea trawl fisheries, Etmopterus lucifer, alongside the Etmopterus granulosus, constitutes the predominant by-catch species. == Taxonomy ==
Taxonomy
Etmopterus lucifer was first discovered and scientifically documented by David Starr Jordan & John Otterbein Snyder in 1902 off the east coast of Honshu, Japan. It is a member of the family Etmopteridae which is in the order Squaliformes. == Anatomy and morphology ==
Anatomy and morphology
Etmopterus lucifer has a slender body with brown coloration on the top and black coloration on the bottom. E. lucifer is shown to have a higher amount of poresassociated with sensory ampullae of Lorenzinion its ventral side than dorsal side, meaning the species likely closes in on prey from above. Black markings are also featured around the pelvic, or ventral, fins and at the base of the caudal, or tail, fin. The blackbelly lanternshark has two dorsal fins, the second of which is larger, and a caudal, or tail, fin, which is relatively long. E. lucifer has relatively long gills, as well as five branchial arches. Teeth are around 2 mm in length. Photophores are located across the body of E. lucifer in distinct arrangements, but are found in much higher density on the ventral side of the shark where they are oriented downward. E. lucifer also displays photophores in clade specific lateral markings as well as along its pectoral fins and claspers. Bioluminescence in blackbelly lanternsharks and other Etmopteridae sharks has been found to be controllable through manipulation of hormones including melatonin (MT), α-melanocyte-stimulating hormone (α-MSH), and adrenocorticotropic hormone (ACTH). MT was found to induce light emission in the photophores of E. lucifer, a process that simultaneously occurred with the opening of the iris-like structure (ILS) leading researchers to conclude that both the bioluminescent chemical reaction occurring in photocytes and the ILS appear to be involved in the control of light emissions. α-MSH and ACTH were both found to decrease light emissions in the photophores of E. lucifer. Additional information on the biochemical mechanism for bioluminescence in sharks is still being researched. Studies have attempted to link bioluminescence to known luciferins, chemicals that produced light when oxidized by a luciferase enzyme, but it has been suggested that a novel photoprotein or luciferase might be present in sharks. == Biology and ecology ==
Biology and ecology
Blackbelly lanternsharks are presumed to be ovoviviparous. (lanternfishes) with slight regional differences. Populations found near Australia have been found to consume a variety of teleost, with the most common being myctophids, while also being known to feed on crustaceans and squids. == Conservation status ==
Conservation status
In June 2018, the New Zealand Department of Conservation classified the blackbelly lanternshark as "Not Threatened" with the qualifiers "Data Poor" and "Secure Overseas" under the New Zealand Threat Classification System. ==References==
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