Sister groups to Ferae According to recent studies, the closest relatives of Ferae are members of clade
Pan-Euungulata (group that includes mirorder
Euungulata and extinct genus
Protungulatum). Together they form grandorder
Ferungulata. An alternate phylogeny holds that the closest relative to Ferae is order
Perissodactyla, with whom they form a clade
Zooamata. Together, clade Zooamata and order
Chiroptera form clade
Pegasoferae, and Pegasoferae is sister taxon to order
Artiodactyla within clade
Scrotifera. However, subsequent molecular studies have generally failed to support this proposal.
Position of pangolins and creodonts within clade Pangolins were long thought to be the closest relatives of
aardvark and
xenarthrans, forming to the now obsolete order
Edentata. Research based on immunodiffusion technique and comparison of protein and DNA sequences revealed the close relationships between pangolins and
carnivorans, with whom they also share a few unusual derived morphological and anatomical traits, such as the ossified
tentorium cerebelli and the fusion of the
scaphoid and
lunate bones in the wrist. The
last common ancestor of extant Ferae is supposed to have diversified c. 79.47 million years ago. and
Altacreodus.
Taxonomy Alternative classification and possible fossil members In
Halliday et al. (2015) various enigmatic Palaeocene
eutherian mammals have been proposed to be possible members of Ferae, like members of orders
Mesonychia,
Pantodonta and
Taeniodonta, and families
Arctocyonidae,
Didelphodontidae,
Nyctitheriidae,
Palaeoryctidae,
Periptychidae and
Triisodontidae. Mesonychians are proposed to be a sister group to carnivoramorphs, while arctocyonids were
polyphyletic, with genera
Arctocyon and
Loxolophus as a sister taxa to pantodonts and periptychids,
Goniacodon and
Eoconodon sister to the Carnivoramorpha-Mesonychia clade, and other genera allied with creodonts and palaeoryctids. == See also ==