The exact way in which pastoralism reached East Africa during the Pastoral Neolithic is not completely understood. The pottery and stone tools found near
Lake Turkana supports that migrants from Ethiopia and Sudan traveled south in small bursts and introduced pastoralism. A considerable amount of evidence supports the case of there being two major expansions (associated with the spread of
Afro-Asiatic and
Nilo-Saharan languages) in eastern Africa which transformed the food systems of the region. A study by Prendergast et al. (2019) analysed genome-wide DNA data from 31 Pastoral Neolithic individuals from sites in Kenya and Tanzania. The study found that these early pastoralists harboured ancestry from three distinct ancient populations, related to: (1) modern groups from northern Africa and the
Levant, (2) contemporary
Nilotic speakers such as the Dinka or Nuer, and (3) hunter-gatherers from East Africa. The Pastoral Neolithic individuals were modelled as deriving ~40% of their ancestry from Chalcolithic Levantines (sampled by Harney et al. 2018), ~40% from a population related to present-day Dinka, and ~20% from East African hunter-gatherers, represented by an ancient forager from
Mota in Ethiopia. A study by Skoglund et al. (2017) similarly found that a Pastoral Neolithic individual from Tanzania, dating from ~3,100 BP, derived ~38% of her ancestry from Neolithic farmers of the Levant. According to the authors, this result could be explained by "the migration into Africa by descendants of
pre-pottery Levantine farmers." A study by Wang et al. (2022) analyzed a sample from Kadruka in Upper Nubia, dated to roughly 4000 BP (c. 2000 BC), and found it to be genetically indistinguishable from those of the Pastoral Neolithic, harbouring a similar mix of approximately 40% Levantine-related and 60% East African-related ancestry. The Kadruka individual was from an agro-pastoral population linked with the
Kerma culture of Upper Nubia. A study by Vicente et al. (2021) found that people related to the Pastoral Neolithic introduced pastoralism into southern Africa approximately 2000 years ago, and admixed with local hunter-gatherer groups to form the
Khoekhoe populations in South Africa. These pastoralists carried approximately 69% East African and 31% Eurasian ancestry, and were likely Cushitic speakers. Lin et al. (2018) found that this pastoralist migration introduced the
SLC24A5 Eurasian light-skin pigmentation gene into the
Khoe-San population 2,000 years ago, which subsequently experienced a
selective sweep within the Khoe-San. The most common SLC24A5
haplotype was found to be identical among European, eastern African, and Khoe-San individuals. All extant Khoe-San groups have admixture with a mixed group containing East African and Eurasian ancestry. Marshall et al (2002) take the tenth millennium BP as that of African cattle domestication. Plant domestication is placed by these researchers as being sometime after 4000 BP. Their point is that the advantage of yield is not, in the African context, a significant driver compared to the risks of aridity and the need to move so as to ensure feed for cattle. These are conclusions based on remains. == Cultural characteristics ==