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Paul Portier (physiologist)

Paul Jules Portier was a French physiologist who made important contributions to the discovery of anaphylaxis and the development of symbiogenesis. On a scientific expedition organised by Albert I, Prince of Monaco, he and Charles Richet discovered that toxins produced by marine animals could induce fatal shocks. They named the medical phenomenon "anaphylaxis," from which Richet went on to receive the 1913 Nobel Prize in Physiology or Medicine. Portier was the first scientist to explain that the cell organelle, mitochondrion, arose by symbiosis according to his evolutionary theory in 1918.

Biography
Portier was born in Bar-sur-Seine, France, to Ernest Paul and Julie Moreau Laure. He studied elementary education at the Lycée de Troyes from 1878 to 1885. After passing the final secondary examination (called bac) from the Saint-Sigisbert in Nancy, he qualified for service in the Ministry of Finance in 1888. However, he chose to study biology, following his childhood dream. In 1889, he entered the University of Paris from where he earned an M.D. in 1897 and Doctor of Science (docteur ès sciences) degree in 1912. In 1920, he was appointed professor of professor of comparative physiology at the University of Paris. In 1923, the University of Paris created a chair of physiology, which he held for the rest of his career. He retired in 1936, and the university awarded him the position of honorary professor. He played active roles in the administrations of the French Academy of Sciences and the French Academy of Medicine. He published his last book The Biology of Butterflies in 1949. Portier married Françoise Noiret Claudine in 1911, and had three daughters, Andrée, Jeannine and Paulette. He spent his last days at his home in Bourg-la-Reine. == Contributions ==
Contributions
Anaphylaxis In 1901, Albert I, Prince of Monaco organised a scientific expedition around the French coast of the Atlantic Ocean. He specifically invited Portier and Charles Richet, professor of physiology at the Collège de France, to join him for investigating the toxins produced by cnidarians (like jellyfish and sea anemones). Richet and Portier boarded Albert's ship Princesse Alice II from where they collected various marine animals. Richet and Portier extracted a toxin called hypnotoxin from their collection of jellyfish (but the real source was later identified as Portuguese man o' war) and sea anemone (Actinia sulcata) from Cape Verde Islands. In their first experiment on the ship, they injected a dog with the toxin in an attempt to immunise the dog, which instead developed a severe reaction (hypersensitivity). To confirm the findings, they knew that more experimental works were needed in the laboratory. In 1902, Richet introduced the term aphylaxis to describe the condition of lack of protection. He later changed the term to anaphylaxis on grounds of euphony. The term is from the Greek ἀνά-, ana-, meaning "against", and φύλαξις, phylaxis, meaning "protection". On 15 February 1902, Richet and Portier jointly presented their findings before the Société de biologie in Paris. The moment is regarded as the birth of allergy (the term invented by Clemens von Pirquet in 1906) study (allergology). Portier never claimed the co-discovery of anaphylaxis, instead honoured Richet as a senior scientist. After the Nobel Prize, Richet praised him for "giving up all claim to the honor of the discovery." It was at the time a known fact that such bacteria were parasites. Portier began to realised that microbes could be necessary for the lives and formation of higher organisms. In 1917, he published the role of symbiosis in the lives of plants and animals, and by that time he started writing a book, he called Les Symbiotes. In 1918, Portier, summing up his observations on symbiosis in nature and his evolutionary idea (now known as symbiogenesis), published Les Symbiotes, dedicating it to Prince Albert.As Portier himself remarked that his theory was "a veritable scientific heresy," The next year, Auguste Lumière published a refutation Le Mythe des Symbiotes ("The Myth of Symbiotes").) == Honours ==
Honours
Portier received the Montyon Prize in 1912, La Caze in 1934, and Jean Toy in 1951 from the French Academy of Sciences. He was given Chevalier (Knight) in 1923, Officier (Officer) in 1935, and Commandeur (Commander) in 1951 of the Legion of Honour. He was honoured Commander of the Order of Saint Charles (1951) and of the Order of Cultural Merit (1954) of Monaco. He was elected member of the French Academy of Medicine in 1929 and of the French Academy of Sciences in 1936. == References ==
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