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Cycas

Cycas is a genus of cycad, and the only genus in the family Cycadaceae with all other genera of cycad being within the Zamiaceae family. Cycas circinalis, a species endemic to India, was the first cycad species to be described in western literature, and is the type species of the genus.

Range
The genus Cycas is native to parts of Asia, eastern Africa and Oceania. In Asia, Cycas (and therefore, the family Cycadaceae) represent the only cycads native to Asia. Within Asia, Cycas species are native from India and Sri Lanka in the west, through China to Japan in the north east and through south east Asia (including the Philippines) to Indonesia in the south. Globally, the northernmost species ( Cycas revoluta) is found at 31°N in southern Japan. In Oceania, Cycas species are native to Australia, Papua New Guinea, and the island nations of the Pacific Ocean region, but are absent from New Zealand. As in Asia, Cycas is the only genus of cycad found across the region, with the exception for Australia, where cycads native to all three families of cycads are found. Despite this, Australia also has the largest number of Cycas species globally with 34 native Cycas species accepted (as of April 2024), including the southernmost species globally (Cycas megacarpa) found at 26°S in southeast Queensland. At least 7 Cycas species are found in Papua New Guinea, some of which are also found in parts of neighbouring Indonesia. Globally, some Cycas species are considered to be relatively widespread, for example Cycas thouarsii is native to a large area of mainland Africa and islands of the Indian Ocean, and Cycas pectinata and Cycas clivicola are both native to large areas of Asia; however, most Cycas species have restricted distributions, with some restricted to very small areas. In Asia, for example, more than 75% of species occur in no more than one country. For some countries, even though the number of species may not be high, the level of species-specific genetic variation can be very high meaning that so that even countries with few species, it may contain distinct gene pools of widespread species. ==Evolution==
Evolution
Cycas (and the Cycadaceae family) is considered as being an early offshoot that diverged early from all other living cycads (which belong to the family Zamiaceae). Estimates of the timing of the split ranging from the Jurassic to the Carboniferous. The presence of Cycas in Australasia and eastern Africa is relatively new, but the major evolutionary events behind the genesis of new species have taken place in Indochina and Australia, where the majority of living species are native.' A later study suggested that these seeds could not be assigned to the stem-group of Cycas with confidence due to lacking the double vascular system that characterises the seeds of all living cycads. The leaf fossil genus Paracycas known from the Jurassic and Cretaceous of Europe has been suggested to be early representatives of the Cycas'' lineage by cladistic analysis. '' ==Conservation status==
Conservation status
Cycas species are threatened worldwide and almost all the species are listed in the IUCN Red List. Cycas beddomei is the only species of the genus Cycas listed in Appendix I of CITES. All other members of Cycadaceae are listed under Appendix II. Cycas rumphii and Cycas pectinata have the most widespread distribution. ==Phylogeny==
Phylogeny
'' megasporophylls with nearly-mature seeds on a wild plant in north Queensland, Australia '' in north Queensland '' in north Queensland with new flush of fronds during the rainy season, still with glaucous bloom Other species: • Cycas aenigma K.D.Hill & A.Lindstr. • Cycas apoa K.D.Hill • Cycas arnhemica K.D.Hill • Cycas badensis K.D.Hill • Cycas bglswamyi Pant ex Agrawal, Akhtar & R.C.Srivast. • Cycas brunnea K.D.Hill • Cycas campestris K.D.Hill • Cycas candida K.D.Hill • Cycas cantafolia Jutta, K.L.Chew & Saw • Cycas changjiangensis N.Liu • Cycas chenii X.Gong & Wei Zhou • Cycas collina K.D.Hill, H.T.Nguyen & P.K.Lôc • Cycas cupida P.I.Forst. • Cycas darshii R.C.Srivast. & Jana • Cycas desolata P.I.Forst. • Cycas dharmrajii L.J.Singh • Cycas distans P.I.Forst. & B.Gray • Cycas divyadarshanii Khuraijam & Rita Singh • Cycas elephantipes A.Lindstr. & K.D.Hill • Cycas falcata K.D.Hill • Cycas flabellata Agoo, Madulid & J.R.Callado • Cycas fugax K.D.Hill, H.T.Nguyen & P.K.Lôc • Cycas furfuracea W.Fitzg. • Cycas glauca Miq. • Cycas hoabinhensis Lôc & H.T.Nguyen • Cycas inermis Oudem. • Cycas javana (Miq.) de Laub. • Cycas lacrimans A.Lindstr. & K.D.Hill • Cycas lane-poolei C.A.Gardner • Cycas laotica T.H.Nguyên & K.S.Nguyen • Cycas litoralis K.D.Hill • Cycas mindanaensis Agoo, Madulid & J.R.Callado • Cycas miquelii Warb. • Cycas montana A.Lindstr. & K.D.Hill • Cycas nayagarhensis Rita Singh, P.Radha & Khuraijam • Cycas nathorstii J.Schust. • Cycas orixensis (Haines) Rita Singh & Khuraijam • Cycas pachypoda K.D.Hill • Cycas panzhihuaensis Zhou & S.Y.Yang • Cycas papuana F.Muell. • Cycas pectinataCycas pranburiensis S.L.Yang et al. • Cycas pruinosa Maconochie • Cycas sainathii R.C.Srivast. • Cycas sancti-lasallei Agoo & Madulid • Cycas saxatilis K.D.Hill & A.Lindstr. • Cycas scratchleyana F.Muell. • Cycas seemannii A.Braun • Cycas seshachalamensis P.V.C.Rao et al. • Cycas sexseminifera F.N.Wei • Cycas shanyaensis Fu • Cycas sundaica Miq. ex A.Lindstr. & K.D.Hill • Cycas terryana P.I.Forst. • Cycas tuckeri K.D.Hill • Cycas xipholepis K.D.Hill • Cycas yorkiana K.D.Hill • Cycas yunnanensis Cheng, Fu & Cheng • Cycas zambalensis Madulid & Agoo • Cycas zeylanica (J.Schust.) A.Lindstr. & K.D.Hill ==References==
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