The pathogen is a
macrocyclic,
autoecious rust fungus, and produces five different
spore states that represent the asexual and sexual components of the life cycle.
Dikaryotic urediniospores are released during the summer as well as the spring while
teliospores represent the
overwintering stage. When the sexual outcrossing occurs the dikaryotic
aeciospores and urediniospores are initiated. The teliospores in the spring will then undergo
meiosis in order to produce
haploid basidiospores. This follows the insect-mediated transfer of
spermatia from the
spermogonia of the different types of mating to the receptive
hyphae. The spores are spread very easily and can be spread by wind. ==Management==