Pinus heldreichii

It is an evergreen tree up to in height, and in trunk diameter. The bark is smooth pale grey at first, becoming hard and tesselated into small hexagonal to rounded plates (somewhat reminiscent of crocodile skin) in old trees. It is a member of the hard pine group, Pinus subgenus Pinus, with leaves ('needles') in fascicles (bundles) of two, with a persistent sheath. They are long and thick. Cones are long, with thin, fragile scales; they are dark blue-purple before maturation, turning brown when ripe about 16–18 months after pollination. The long seeds have a wing and are wind-dispersed. File:Байкушева мура 01.JPG|Trunk of an old tree ('Baikushev's pine') File:Pinus heldreichii leg P.Cikovac Jastrebica Mt Orjen.jpg|Bark on an old tree, showing the distinctive "crocodile-skin" pattern File:2024-07-24 Pinus heldreichii, Park Prirode Blidinje, Jablanica, Herzegovina 1.jpg|Dark purple one-year old cones, and a light brown open cone from the previous year File:2024-07-24 Pinus heldreichii, Park Prirode Blidinje, Jablanica, Herzegovina 5.jpg|First-year conelets about two months after pollination File:2024-07-24 Pinus heldreichii, Park Prirode Blidinje, Jablanica, Herzegovina 3.jpg|Mature open cone == Nomenclature ==

The species was first described as Pinus heldreichii by the Swiss botanist K. Hermann Christ in honour of Theodor von Heldreich in 1863 from specimens von Heldreich collected on Mount Olympus, and then described a second time as P. leucodermis in 1864; the author of the second description (the Austrian botanist F. Antoine who found it on Orjen above the Bay of Kotor in Montenegro) being unaware of the slightly earlier publication by Christ. Some minor morphological differences have been claimed between the two descriptions (leading to the maintenance of both as separate taxa by a few botanists), but this is not supported by modern studies of the species, which show that both names refer to the same taxon. The discrepancies in the descriptions are largely due to Christ's cone specimens being immature and shrunken after drying, having been collected in July, three months before maturity. == Taxonomy ==

It can be found in the mountains of Bosnia and Herzegovina, Montenegro, southwestern Bulgaria, Albania, North Macedonia, Kosovo, northern Greece (Valia Kalda, Smolikas and Vasilitsa, Mount Olympus and in other high mountains), and locally in southern Italy (it is the symbol of the Pollino National Park), growing at altitude. It reaches the alpine tree line in these areas. == Longevity ==

A tree in Northern Greece was dated as 1,075 years old in 2016. What is believed to be the oldest known living tree in Europe has been discovered in a remote mountainous area of the Pollino National Park in southern Italy. It is a Heldreich's pine estimated at 1,230 years. Much of its core has turned to dust, but there is enough new growth to confirm it is still alive. A notable specimen in the Pirin Mountains of Bulgaria, known as Baikushev's pine, is tall, in diameter, and is estimated to be over 1300 years old. ==Cultivation and uses==

P. heldreichii is able to adapt to extreme environmental conditions and is also a good colonist of dry high altitude sites. It is resistant to sulphur dioxide, hydrogen fluoride, nitrogen dioxide and ozone pollution and is further able to withstand wind, ice and heavy snow. These abilities makes it suitable for reforestation of extensive dry and high-altitude areas. In the south of Italy it is planted because it is less susceptible to pests than other pine species. Bosnian pine is a popular ornamental tree in parks and large gardens, giving reliable, steady, though not fast, growth on a wide range of sites, and with a very neat, conical crown. It is also noted for its very decorative purple cones. The cultivars 'Smidtii' and 'Compact Gem' have been given the Royal Horticultural Society's Award of Garden Merit. It is hardy down to at least , and tolerant of severe wind exposure. Many in cultivation are still grown under the name Pinus leucodermis or Pinus heldreichii var. leucodermis. The wood of the species is aromatic, containing several types of monoterpenes and sesquiterpenes (essential oils), noticeably limonene and cembrene, and thus is commonly utilised in the Balkans for making wine barrels for white wines. ==Gallery==

File:2024-07-24 Pinus heldreichii, Park Prirode Blidinje, Jablanica, Herzegovina 2.jpg|Young tree File:2024-07-24 Pinus heldreichii, Park Prirode Blidinje, Jablanica, Herzegovina 6.jpg|Smooth bark on a young tree Pinus heldreichii Orjen.jpg|Specimen on Orjen Heldreichii Orjen.jpg|Specimen on Orjen Heldreichii Orjen Beechforest.jpg ==References==

• Simone Morris (2017) Pini Loricati nella nebbia | • Simone Morris (2018) Loricati in the fog | ==External links==