Forming the first evidence of crown-group gastropods when they appeared in the
Upper Cambrian, the fossil record of the Pleurotomarioideans has no substantial gaps until today. The group took quite a hit at the
Cretaceous–Paleogene boundary (K–T boundary), with only the Pleurotomariidae surviving the
Cretaceous–Paleogene extinction event – and then only in deep waters. Living representatives of the group were first discovered in the mid-19th century, and their unusual mix of primitive and derived characters perplexed biologists. The researchers originally responded by re-working their ideas of how the gastropod lineage evolved, but with the introduction of
cladistics, attempts are currently underway to fit them into a molluscan phylogeny. ==Taxonomy ==