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Postorbital bar

The postorbital bar is a bony arched structure that connects the frontal bone of the skull to the zygomatic arch, which runs laterally around the eye socket. It is a trait that occurs in some mammalian taxa, such as most strepsirrhine primates and the hyrax, while haplorhine primates have evolved fully enclosed sockets. One theory for this evolutionary difference is the relative importance of vision to both orders. As haplorrhines tend to be diurnal, and rely heavily on visual input, many strepsirrhines are nocturnal and have a decreased reliance on visual input.

Function
In the past decades, many different hypotheses were made on the possible function of the postorbital bar. Three of them are commonly cited. External trauma hypothesis Prince and Simons offered the external trauma hypothesis, where the postorbital bar protects the orbital contents from external trauma. However, a few years later Cartmill offered a new view on this bone and came up with the mastication hypothesis. Greaves suggests that the bar strengthens the relatively weak orbital area against torsional loading, imposed by bite force in species with large masseter and temporalis muscles. However the orientation of the postorbital process does not match the direction of the forces mentioned by Greaves. Position hypothesis Cartmill suggests that in small mammals with large eyes and relatively small temporal fossae, where the anterior temporal muscle and the temporalis fascia are pulled to a more lateral position with increasing orbital convergence (front-facing eyes), the tension caused by the contraction of these muscles would distort the orbital margins and disrupt oculomotor precision. Heesy shows that the postorbital bar stiffens the lateral orbit. Without a stiffened lateral orbit, deformation would displace soft tissues, when contraction of the anterior temporalis muscle takes place, thus impeding eye movement. == Occurrence ==
Occurrence
A complete postorbital bar has evolved at least eleven times as a convergent adaptation in nine mammalian orders. == Postorbital process ==
Postorbital process
Postorbital bars are likely derived from well-developed postorbital processes, an intermediate condition where a small gap retains between the process and the zygomatic arch. Well-developed postorbital processes have evolved separately within the orders of the Dermoptera and Hyracoidae and the Chiropteran families of Emballonuridae and Pteropodidae and to varying degrees within many carnivorian taxa. ==References==
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