, combining its scarce remains with a now outdated assumed affinity with the macraucheniid
Theosodon. Protheosodon coniferus was first described in 1897 by
Florentino Ameghino, based on fossils found in
Deseadan deposits from
Chubut Province,
Argentina. Other fossils attributed to the genus have been found in slightly older deposits in
Colombia.
Protheosodon was initially thought to be a member of
Macraucheniidae, mainly based on characteristics from its lower teeth. Subsequent researches has indicated that it was more probably nested within
Proterotheriidae, a group of litopterns which, in the course of their evolution, developed horse-like forms, well adapted for running. It may have been a member of
Anisolambdinae, the most basal subfamily of proterotheriids.
Protheosodon seems to have been part of a collateral branch, contemporary to other, more derived, proterotheriids. ==Bibliography==