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Bagworm moth

The Psychidae are a family of the Lepidoptera. The bagworm family is fairly small, with about 1,350 species described. Bagworm species are found globally, with some, such as the snailcase bagworm, in modern times settling continents where they are not native.

Description
The caterpillar larvae of Psychidae construct cases out of silk and environmental materials such as sand, soil, lichen, or plant materials. These cases are attached to rocks, trees or fences while resting or during their pupa stage, but are otherwise mobile. The larvae of some species eat lichen, while others prefer green leaves. In many species, the adult females lack wings and are therefore difficult to identify accurately. Case-bearer cases are usually much smaller, flimsier, and consist mainly of silk, while bagworm "bags" resemble caddisfly cases in their outward appearance – a mass of (mainly) plant detritus spun together with silk on the inside. Bagworm cases range in size from less than 1 cm to 15 cm among some tropical species. Each species makes a case particular to its species, making the case more useful to identify the species than the creature itself. Cases among the more primitive species are flat. More specialized species exhibit a greater variety of case size, shape, and composition, usually narrowing on both ends. The attachment substance used to affix the bag to a host plant or structure has a higher tensile strength than that of other moth species, requiring greater force to initiate breakage. Body markings are rare. Adult females of many bagworm species are larviform, with only vestigial wings, legs, and mouthparts. In some species, parthenogenesis is known. The adult males of most species are strong fliers with well-developed wings and feathery antennae but survive only long enough to reproduce due to underdeveloped mouthparts that prevent them from feeding. Most male bagworm wings have few of the scales characteristic of most moths, instead having a thin covering of hairs. File:Pachythelia villosella.case.jpg|"Bag" of Pachythelia villosella (Oiketicinae) File:Grasninella m.JPG|Adult specimen of Phalacropterix graslinella (Oiketicinae) File:Bagworm Moth larva Negev.JPG|Bagworm larva in the Negev (April 2014). Case is made mostly of feathery stork's bill seeds (Erodium cicutarium). File:Family Psychidae444.jpg|Bagworm (possibly Hyalarcta huebneri) extending its forequarters from its case in the act of locomotion. ==Ecology==
Ecology
In the larval stage, bagworms extend their head and thorax from their mobile case to devour the leaves of host plants, often leading to the defoliation of their hosts. Trees infested with bagworms exhibit increasingly damaged foliage as the infestation increases, which may result in gradual mortality accrual, especially on tree plantations in tropical regions where bagworms are more frequently multivoltine. Some bagworms are specialized in their host plants (monophagous), while others can feed on a variety of plant species (polyphagous). A few species also consume small arthropods (such as the camphor scale Pseudaonidia duplex, a scale insect). One bagworm species was found to eat an orb-web of Plebs sachalinensis (Araneae, Araneidae) entirely. Since bagworm cases are composed of silk and the materials from their habitat, they are naturally camouflaged from predators. Predators include birds and other insects. Birds often eat the egg-laden bodies of female bagworms after they have died. Since the eggs are very hard-shelled, they can pass through the bird's digestive system unharmed, promoting the spread of the species over wide areas. A bagworm begins to build its case as soon as it hatches. Once the case is built, only adult males ever leave the case, never to return, when they take flight to find a mate. Bagworms add material to the front of the case as they grow, excreting waste materials through the opening in the back of the case. When satiated with leaves, a bagworm caterpillar secures its case and pupates. The adult female, which is wingless, either emerges from the case long enough for breeding or remains in the case while the male extends his abdomen into the female's case to breed. Females lay their eggs in their case and die. The female evergreen bagworm (Thyridopteryx ephemeraeformis) dies without laying eggs, and the larval bagworm offspring emerge from the parent's body. Some bagworm species are parthenogenetic, meaning their eggs develop without male fertilization. Each bagworm generation lives just long enough as adults to mate and reproduce in their annual cycle. == Systematics ==
Systematics
Ten subfamilies and about 240 genera are recognized among the bagworms. The subfamilies of Psychidae, with some notable genera and species also listed, are: Subfamily EpichnopteriginaeAcentraBijugisEpichnopterixEpichnopterix plumellaHeliopsychideaMauropterixMontanimaPsychideaPsychocentraRebeliaReisseroniaStichobasisStichobasis postmeridianusWhittleiaWhittleia retiella Subfamily NaryciinaeDahlica Enderlein, 1912 • Dahlica triquetrellaEosolenobia Filipjev, 1924 • NaryciaPostsolenobia Meier, 1958 • Siederia Subfamily OiketicinaeApteronaApterona helicoidella – (snailcase bagworm) • CanephoraCanephora hirsutaCryptothelea (= Platoeceticus) • Cryptothelea gloveriiAstalaKotochaliaKotochalia junodi – (wattle bagworm) • HyalarctaDeborreaDeborrea malgassa – (fangalabola) • EumetaEumeta crameri – (faggot worm) • MegalophanesMegalophanes viciellaOiketicusZamopsycheEucoloneuraPachytheliaPhalacropterixPtilocephalaPtilocephala plumiferaSterrhopterixSterrhopterix fuscaThyridopteryx Subfamily PlacodominaePlacodoma Subfamily PsychinaeLiothulaLiothula omnivoraLuffiaLuffia ferchaultellaLuffia lapidellaPsychePsyche castaPsyche crassiorellaPsyche rasseiPsyche saxicolellaProchalia Subfamily TaleporiinaeBankesia Tutt, 1899 • Bankesia conspurcatellaBrevantennia Sieder, 1953 • CebysaCebysa leucotelus – (Australian bagmoth) • DiplodomaDiplodoma adspersellaEotaleporia Sauter, 1986 • Praesolenobia Sieder, 1954 • Pseudobankesia Meier, 1963 • Sciopetris Meyrick, 1891 • Taleporia Subfamily TyphoniinaeTyphoniaTyphonia animosaTyphonia bimaculata Incertae sedisEumasiaEumasia crisostomellaEumasia parietariella (Heydenreich, 1851) • IphiergaIphierga chrysophaes Turner, 1917 Subfamily ScoriodytinaeScoriodyta Meyrick, 1888 • Scoriodyta conisalia Meyrick, 1888 Subfamily MetisinaeMetisa Hampson, 1895 • Metisa canifrons Hampson, 1895 Subfamily PseudarbelinaePseudarbela Sauber, 1902 • Pseudarbela celaena (Bethune-Baker, 1904) • Pseudarbela aurea (Bethune-Baker, 1904) • Pseudarbela papuana Clench, 1959 • Pseudarbela semperi Sauber, 1902 • Casana Walker, 1865 • Casana trochiloides Walker, 1865 • Linggana Roepke, 1957 • Linggana cardinaali Roepke, 1957 ==Applications==
Applications
The silk of bagworm moth larvae is reportedly "more durable than spider silk". Japanese pharmaceutical company Kowa and the National Agriculture and Food Research Organization announced a new textile, Minolon, in 2024, that combined carbon fibers with sheets of bagworm silk. The resulting material was claimed to be environmentally friendly, with Kowa planning to sell it to aircraft, automobile, and sporting good manufacturers. ==References==
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