There are currently 14 species in this genus (others are considered
synonyms): (
Gymnura altavela)|thumb •
Gymnura afuerae (Hildebrand, 1946) •
Gymnura altavela (
Linnaeus,
1758) – Spiny butterfly ray •
Gymnura australis (
E. P. Ramsay &
Ogilby, 1886) – Australian butterfly ray •
Gymnura crebripunctata (
W. K. H. Peters, 1869) – Longsnout butterfly ray •
Gymnura hirundo (Lowe, 1843) – Madeira butterfly ray •
Gymnura japonica (
Temminck &
Schlegel, 1850) – Japanese butterfly ray •
Gymnura lessae (
Yokota &
Carvalho, 2017) – Lessa's butterfly ray •
Gymnura marmorata (
J. G. Cooper, 1864) – California butterfly ray •
Gymnura micrura (
Bloch &
J. G. Schneider, 1801) – Smooth butterfly ray •
Gymnura natalensis (
Gilchrist &
W. W. Thompson, 1911) – Backwater butterfly ray •
Gymnura poecilura (
G. Shaw, 1804) – Longtail butterfly ray •
Gymnura sereti (
Yokota &
Carvalho, 2017) – Seret's butterfly ray •
Gymnura tentaculata (
J. P. Müller &
Henle, 1841) – Tentacled butterfly ray •
Gymnura zonura (
Bleeker, 1852) – Zonetail butterfly ray ;Synonyms •
Gymnura bimaculata (Norman, 1925) - synonym of
Gymnura japonica •
Gymnura crooki Fowler, 1934 - synonym of
Gymnura poecilura Fossil taxa The following fossil gymnurid genera are known: • †
Subathunura Kumar & Loyal, 1987 (Eocene of India) • †
Jacquhermania Cappetta, 1982 (Middle to Late Eocene of Europe and eastern North America)'''' • †
Ouledia Cappetta, 1986 (Late Paleocene to Middle Eocene of North Africa, South America and South Asia) • †
Pachygymnura Adnet
et al., 2020 (Late Eocene of Egypt) The following fossil species of
Gymnura are also known: • †
Gymnura delpiti Cappetta, 1984 (
Late Paleocene of Morocco) • †
Gymnura grootaerti Herman, 1984 (
Early Eocene of Belgium) • †
Gymnura hovestadti Herman, 1984 (
Early Oligocene of Belgium) • †
Gymnura laterialata Werner, 1989 (
Cenomanian of Egypt) • †
Gymnura transita Averianov & Udovitshenko, 1993 (Early Eocene of
Kyrgyzstan) ==Diet==