The species was first described in 1860 by
Charles Knight, using the name
Verrucaria moniliformis. It was moved to the genus
Pyrenula in 1894 by
Johannes Müller Argoviensis. The species was only known from its type collection until 2025, when new specimens were discovered in the
Unitec Institute of Technology herbarium, including student specimen collections dating back to 2015. After rediscovery, the species was redescribed in 2025 by Andrew J. Marshall,
Peter de Lange, Dan Blanchon and
André Aptroot, based on fresh specimens. The holotype is held by the
British Museum, and while Knight did not specify a location, is presumed to have been collected from the Auckland Region. == Ecology ==