The Quaternary Period is conventionally divided into two epochs: the Pleistocene , beginning about 2.58 million years ago, and the Holocene , which began about 11,700 years ago. The term Quaternary was first used by Italian engineer
Giovanni Arduino in the late eighteenth century to describe the four most recent geologic eras. It is subsequently regarded as "a phase of highly variable climates, with marked periods of time when global temperatures were significantly lower than today and evidence for which was interpreted by
Louis Agassiz as indications of a geologically recent 'Great Ice Age'". In the early nineteenth century French scientist Georges Cuvier proposed that some animals that lived in the Pleistocene epoch were
made extinct by some environmental 'revolution' (e.g. some catastrophic flooding events). In the twentieth century
Milutin Milankovitch, a Serbian mathematician and
geophysicist, developed a theory linking long-term climate changes to the motion of the Earth. It offered information about the changes in seasonal "
insolation" (incident solar radiation) over millions of years.
André Berger, a Belgian
climatologist, identified certain time periods where reconstructed insolation deviated significantly from the average. Many of his analyses show that from May to August, there has been a forwarded shift of insolation maximum (higher than average) during the late Quaternary. This feature is known as "insolation signature" and may relate to changes in climate as contemplated by Berger. Subsequent work in
palaeoecology,
palaeontology and
palaeoclimatology has revealed relationships between changes in the environment and the planet's history during the Quaternary. Quaternary science has also yielded insight into human
colonization and mobility, providing data about the environment and landscape affecting
human evolution. ==Biosphere impact==