Nuclear radiation is harmful to the environment over immediate (seconds or fractions thereof) as well as long-term (years or centuries) timescales, and it affects the environment on both microscopic (
DNA) and macroscopic (population) levels. Degrees of these effects are dependent on external factors, especially in the case of humans. Radioecology encompasses all radiological interactions affecting biological and geological material as well as those between different phases of matter, as each is capable of carrying radionuclides. Occasionally, the origin of
radionuclides in the environment is actually nature itself, as some geological sites are rich in radioactive uranium or produce radon emissions. The largest source, however, is artificial pollution via nuclear meltdowns or expulsion of radioactive waste from industrial plants. The ecosystems at risk may also be fully or partially natural. An example of a fully natural ecosystem might be a
meadow or
old-growth forest affected by fallout from a nuclear accident such as Chernobyl or Fukushima, while a semi-natural ecosystem might be a
secondary forest, farm, reservoir, or fishery that is at risk of infection from some source of radionuclides. Basic
herbaceous or
bivalve species such as
mosses,
lichens,
clams, and
mussels are often the first organisms affected by fallout in an ecosystem, as they are in closest proximity to the abiotic sources of radionuclides (atmospheric, geological, or aquatic transfer). These organisms often possess the highest measurable concentrations of radionuclides, making them ideal bioindicators for sampling radioactivity in ecosystems. In the absence of sufficient data, radioecologists must often rely on analogs of a radionuclide to attempt to evaluate or hypothesize about certain
ecotoxicological or metabolic effects of rarer radionuclides. In general, techniques in radioecology focus on the study of environmental
bioelectromagnetism,
bioelectrochemistry,
electromagnetic pollution, and
isotope analysis. == Radioecological threats ==