It was initially described as a distinct species by
Robert Fisher Tomes in 1858, but later studies reclassified as conspecific with
Hodgson's bat (
M. formosus). However, a 2014 morphological study found major divergence between
M. formosus and
M. rufoniger, and thus split them from one another. Creating a phylogeny based on wing patterns, the study found
M. rufoniger to be most closely related to
Welwitsch's bat (
M. welwitschii) of
Africa, rather than to
M. formosus. This has also been followed by the
American Society of Mammalogists, the
IUCN Red List, and the
ITIS. A 2017 genetic study found
M. rufoniger to have a low
effective population size, being the lowest of the four
Myotis species sampled for the study. The study found its effective population size to have peaked during the late
Pleistocene, about 50,000 years ago, and to have dramatically decreased since the
Last Glacial Maximum, reaching its lowest level in the present day. == Distribution and habitat ==