SGA has been best studied in
pigs, both due to industry pressures of high mortality rates of
preweaning piglets and the use of pig as a
model organism in science. Runts are caused by interplay between genetics, environment in utero, maternal environment and care. Breeding for larger litter sizes has resulted in there being born more piglets than the
teats of the sow, longer birthing times and more
hypoxic young. Further causes of fetal malfunctioning can be a
circovirus infection, maternal malnutrition or a small or inconveniently placed placenta. In cattle and sheep, an additional reason may be
hot weather during pregnancy. In dairy cows, a contributing factor may be lactating while pregnant, which can overtax the cow's ability to provide sufficient nutrients to the fetus.
Nulliparous cows are more at risk of giving birth to SGA calves, and on average give birth to calves of a lower birth weight. In
dogs, a larger litter size may cause more low weight puppies to be born. Typically low weight puppies, like piglets, have smaller
placentas in comparison to their normal body weight littermates. In
cats, younger mothers are more likely to give birth to kittens with lower body weight. Likelihood of giving birth to low body weight kittens increases if there is at least one
stillbirth in the litter. ==Management==