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Scopolia

Scopolia is a genus of four species of flowering plants in the family Solanaceae, native to Europe and Asia. The genus is named after Giovanni Scopoli (1723–88), a Tyrolean naturalist. The genus has a disjunct distribution, with two recognised species in Central to Eastern Europe,, and two species in East Asia. The two European species are:Scopolia carniolica Jacq. of Slovenia, Austria and the Carpathian Mountains Scopolia caucasica Kolesn. ex Kreyer of the Caucasus

Comparison of Scopolia carniolica with Atropa belladonna
'', showing hairy, lobed calyx and hairy, purple and green, urn-shaped corolla, netted with veins. Note curved pistil (bearing minutely hairy stigma) protruding beyond corolla. The existence of the synonym Scopolia atropoides (i.e. "Atropa-like Scopolia") for Scopolia carniolica demonstrates the perceived similarity between Scopolia carniolica and its better-known relative Deadly Nightshade (Atropa belladonna). The most obvious dissimilarity lies in the respective fruits, that of Scopolia being a pyxidium (i.e. a dry, pot-like capsule with an operculum (lid)) while that of Atropa is a juicy, glistening, jet-black berry bearing a superficial resemblance to a cherry - indeed this pyxidium / berry dichotomy constitutes the feature separating the genus Atropa into a subtribe of its own within the Solanaceous tribe Hyoscyameae: all other genera in tribe Hyoscyameae have the same type of dry, pyxidial capsule as Scopolia. Other points of dissimilarity include: • Pedicel type: Scopolia carniolica, long, slender and delicate / Atropa belladonna, shorter and broadening toward the calyx - particularly at fruiting. • Calyx type: Scopolia carniolica, cup-shaped, with slight peaks rather than distinct lobes / Atropa belladonna distinctly lobed, the individual lobes somewhat leaf-like and these lobes spreading to make the calyx star-shaped in fruit. • Colour of corolla: Scopolia carniolica, exterior: purplish-brown striped longitudinally with cream veins, interior: cream, except for some dark venation at the very base / Atropa belladonna, dull purple shading to green netted with darker veins at the base. • Glabrescence / pubescence: Scopolia carniolica, glabrous (hairless), the corolla often having a glossy, "enameled" appearance / Atropa belladonna pubescent (hairy), the corolla and calyx being clothed in short trichomes (hairs) - though less so in those of its eastern subspecies A. belladonna ssp. caucasica. • Corolla shape: Scopolia carniolica, simple, un-lobed, straight-sided bell / Atropa belladonna urceolate (= urn-shaped) bell bearing short but distinct lobes, somewhat recurved. • Pistil: Scopolia carniolica, straight and shorter than corolla / Atropa belladonna, curved and exserted (=protruding beyond corolla). ==Gallery==
Gallery
File:Scopolia carniolica Finland 2016-04-30.jpg|Scopolia carniolica fresh Spring growth with abundant flowers, old arboretum, Finland. File:Scopolia caucasica 129017727.jpg|Scopolia caucasica in flower, showing distinctive corolla shape - more flared than in other species File:Scopolia japonica-03.jpg|Scopolia japonica (Common name in Japanese: 走りどころ Hashiri-dokoro), plant in flower, Korea. File:Eykman1883-Scopoliae Rhizoma.jpg|Scopoliae Rhizoma (root of Scopolia japonica) ==References==
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