The vestibular system detects linear and angular (rotational) acceleration using specialized organs in the inner ear. Linear accelerations are detected by the
otolith organs, while angular accelerations are detected by the
semicircular canals.
Misleading sensations Without a visual reference or cues, such as a visible horizon, humans will rely on non-visual senses to establish their sense of motion and equilibrium. During the abnormal acceleratory environment of flight, the
vestibular and
proprioceptive systems can be misled, resulting in spatial disorientation. When an aircraft is maneuvering, inertial forces can be created by changes in vehicle speed (linear acceleration) and/or changes in direction (rotational acceleration and
centrifugal force), resulting in perceptual misjudgment of the vertical, as the combined forces of gravity and inertia do not align with what the vestibular system assumes is the vertical direction of gravity (towards the center of the Earth). Under ideal conditions, visual cues will provide sufficient information to override illusory vestibular inputs, but at night or in poor weather, visual inputs can be overwhelmed by these illusory nonvisual sensations, resulting in spatial disorientation. Low visibility flight conditions include night, Similarly, it is possible to gradually climb or descend without a noticeable change in pressure against the seat. In some aircraft, it is possible to execute a loop without pulling negative g-forces so that, without visual reference, the pilot could be upside down without being aware of it. A gradual change in any direction of movement may not be strong enough to activate the vestibular system, so the pilot may not realize that the aircraft is accelerating, decelerating, or banking. , including
attitude indicator (top center) and
turn and slip indicator (bottom left) Gyroscopic
flight instruments such as the
attitude indicator (artificial horizon) and the
turn and slip indicator are designed to provide information to counteract misleading sensations from the non-visual senses.
Otoliths and somatogravic illusions Two otolith organs, the saccule and utricle, are located in each ear and are set at right angles to each other. The utricle detects changes in linear acceleration in the horizontal plane, while the saccule detects linear accelerations in the vertical plane; humans have evolved to assume the vertical acceleration is caused by gravity. However, the saccule and utricle can provide misleading sensory perception when gravity is not limited to the vertical plane, or when vehicle speeds and accelerations result in inertial forces comparable to the force of gravity, as the otoliths only detect acceleration, and cannot distinguish inertial forces from the force of gravity. In a 1954 study (180 – Degree Turn Experiment), the University of Illinois Institute of Aviation found that 19 out of 20 non-instrument-rated subject pilots went into a graveyard spiral soon after entering simulated instrument conditions. The 20th pilot also lost control of his aircraft, but in another maneuver. The average time between onset of instrument conditions and loss of control was 178 seconds. Spatial disorientation can also affect
instrument-rated pilots in certain conditions. A powerful tumbling sensation (
vertigo) can result if the pilot moves his or her head too much during instrument flight. This is called the
Coriolis illusion. Because the semicircular canals are set in three different axes of rotation, if the aviator suddenly moves their head during a rotational acceleration, one canal may abruptly start to detect an angular acceleration while another ceases, resulting in a tumbling sensation. == Visual illusions ==