Below is a
cladogram following a topology by Pêgas and colleagues in 2019. In their analysis, they recovered Targaryendraconia as the sister taxon of the clade
Anhangueria, both of which are within the more inclusive group
Ornithocheirae. Targaryendraconia is split into two families: the
Targaryendraconidae, which contains
Aussiedraco,
Barbosania, and
Targaryendraco, and the
Cimoliopteridae, which contains
Aetodactylus,
Camposipterus, and
Cimoliopterus. }} }} }} }} }} }} }} }} }} Contrastingly, a study by Alexander Averianov in 2025, describing the new targaryendraconian
Saratovia, failed to recover the existence of a cimoliopterid clade. Instead,
Cimoliopterus cuvieri,
Cimoliopterus dunni,
Saratovia, and a grouping of
Camposipterus and
Aetodactylus formed an unresolved
polytomy with Targaryendraconidae. He argued that the distinction between the two groups of targaryendraconian were likely caused due to the incompleteness of known remains; cimoliopterids are characterized by traits of the upper jaw, and targaryendraconids by those of the lower jaw. Most of the known genera are only known from upper or lower jaw, and the complete skull of
Barbosania is preserved in a manner preventing study of the interior surface of the jaw. It was also noted that several uniting characteristics of Anhangueria and Anhangueridae are not evaluable in most or all members of Targaryendraconia. ==References==