Introduction In the introductory chapter, MacArthur and Wilson highlight the significance of studying island biogeography. Since islands are less complex and more numerous than larger ecosystems, islands provide better opportunities to develop insights and perform replicable field research. Given that insular microcosms are common to all ecosystems, principles from island biogeography can be applied generally.
Area-diversity pattern In Chapters 2 and 3, MacArthur and Wilson postulate that insular species richness depends on island size and isolation from source regions. The authors present an equilibrium model that is based on the following concept: when there is an addition of the number of species on an island, the island's immigration rate of new species will decrease while the extinction rate of resident species will increase. MacArthur and Wilson thus assume that there will be an equilibrial point where the immigration rate equals the extinction rate. They further hypothesize that an increase in island size will lower extinction curves while a decrease in distance between the island and the source region will raise immigration curves. Since the intersection of immigration and extinction rate curves determines the species number, the authors predict that larger islands will have more species than smaller islands (assuming these islands are comparably isolated) and isolated islands will have fewer species than islands more proximal to source regions (assuming these islands are equally large). There is additional discourse on how insular clusters and stepping stones affect this model.
Modeling colonization and dispersion Chapter 4 discusses survivorship theory. The authors describe a model which states that the probability for successful colonization is dependent on birth rate, death rate, and carrying capacity of the environment. From this model, conclusions are made on the average survival time for a
propagule's offspring, the average survival time of a saturated population, and characteristics of successful propagules. In Chapter 5, MacArthur and Wilson examine why species can be excluded from insular environments and how the
niche of a species changes after introduction. The authors surmise that pioneering species can be excluded for the following reasons: the island has saturated levels of pre-existing competition, the pioneering species cannot maintain a population large enough to avoid extinction, and the island hosts too many or too few natural predators. When a species colonizes a new area, the authors state that the species will either shift, expand or contract its realized niche. Chapter 6 is a theoretical exploration of dispersal models. The authors consider how insular stepping stones affect the dispersion of species—particularly, the effects that size and isolation of stepping stones have on dispersion. Further consideration is given to how dispersal curves and average distance travelled by pioneers impacts this study. In Chapter 7, the authors state that there are generally three consecutive phases to the evolution of populations after colonization. Initially, there is a trend for colonizers to evolve from
r-strategists into
K-strategists. The
founder effect may also influence colonizing populations during this first phase. The second phase is marked by long term adaptations to the local environment. In this period, abilities for dispersal are commonly reduced, and colonizers will either differentiate or assimilate with competing species. In the third phase, the evolution of colonizing populations may result in
speciation and/or
adaptive radiation. == Field testing ==