Archibald
et al. in 2011 erected the genus
Titanomyrma, described the species
Titanomyrma lubei and proposed two new combinations,
T. gigantea (formerly
Formicium giganteum Lutz, 1986) and
T. simillima (formerly
Formicium simillimum Lutz, 1986).
T. gigantea has been designated the
type species for the genus. The name of the genus is a derivative of the Greek (''''), meaning 'one of prodigious size, strength, or achievement', and alluding to the
Titans of Greek mythology; and the Greek word ('''') meaning 'ant'. The genus
Titanomyrma is differentiated from others in the family by the shape of the
gaster which is variable. In the three included species the gaster ranges from ovate to more slender or cylindrical. The A5 abdominal segment width relative to other gaster segments is variable and the relative lengths of A3–A7 are also variable. near the village of
Messel, in the state of
Hessen, south of
Frankfurt am Main in
Germany.
T. simillima Titanomyrma simillima, as with
T. gigantea, are also only known from the Messel Formation, After discovery, Lube donated the fossil to the collections of the
Denver Museum of Nature and Science where it was noticed by paleoentomologist Bruce Archibald and (then) museum chief curator
Kirk R. Johnson while they were going through storage drawers. Archibald, Mathewes, & Aase (2023) reported a
Titanomyrma queen from the
Allenby Formation, and noted the range extension for
Formiciinae into the
Eocene Okanagan Highlands, as the subfamily was previously considered a strictly
thermophilic ant group. Due to complications arising from preservational distortion during
diagenesis, they were unable to determine the correct size of the queen in life. If the distortion was lateral, then compression to bilateral symmetry yielded an adult length of approximately , placing it the same range as
Formicium berryi and
F. brodiei, known only from wings, and suggested as possible males. Conversely stretching the fossil to bilateral symmetry results in a larger length estimate, placing it as comparable to queens of
T. lubei and
T. simillima. ==Paleoecological implications==