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Erettopterus

Erettopterus is a genus of large predatory eurypterid, an extinct group of aquatic arthropods. Fossils of Erettopterus have been discovered in deposits ranging from Early Silurian to the Early Devonian, and have been referred to several different species. Fossils have been recovered from three continents; Asia, Europe and North America. The genus name is composed by the Ancient Greek words ἐρέττω, which means "rower", and πτερόν, which means "wing", and therefore, "rower wing".

Description
Erettopterus was a big eurypterid, with the largest well-preserved examples of E. osiliensis, the largest species, measuring approximately in length. has been estimated to have reached a length of 1.65 metres (5.4 ft). although the fossils of E. globiceps could be remains of juvenile specimens, which would mean that the species could have reached larger sizes. The telson was expanded and bilobed (divided into two lobes). This form of telson is its main characteristic and differentiates it from the rest of the genera of the family Pterygotidae. Historically it was thought that Erettopterus had five joints in its chelicerae, but studies with complete specimens of Erettopterus and Acutiramus have revealed that the actual count appears to be four joints. The form of the chelicera within Erettopterus was very variable, but they were generally very long with small curved teeth without serrations. Its metastoma (a large plate that is part of the abdomen) was very narrow and cordated (heart-shaped) anteriorly deeply notched (V-shaped slit). The swimming legs were short. The body was elongate-oval in form and gradually attenuated into the abdomen. The compound eyes were broadly crescentic and convex, and as in the rest of the pterygotioids, they were located in the margin of the carapace. ==History of research==
History of research
19th century A total of 19 valid species have been assigned to Erettopterus. Most of the species have been found in the United States or Great Britain, although fossils have also been found in Canada, Scandinavia and Estonia. The type species and the only species described in Scotland, E. bilobus, was first found in Lesmahagow, Scotland, and described by John William Salter in 1856 as a species of Himantopterus (a name that is a junior homonym of a moth genus), and Salter replaced the genus name with Erettopterus in 1859, although it would later be referred to as a species of Pterygotus. It is a very well known and very abundant species that since its original description has not received much attention, so it needs a redescription like other eurypterids to accommode a modern understanding of eurypterid systematics and evolution. The specific name bilobus refers to the bilobed telson of the species. The English geologist Salter recognized in 1859 that it was possible to divide the genus Pterygotus based on the morphology of the telsons of the species that had been assigned to it. He divided Pterygotus into subgenera, including Pterygotus (Erettopterus) for species with a bilobed telson. (eréttō, rower The other was E. grandis (grandis because of the large size of the species) The rami end in a well-developed sharp and triangular point. from the Lochkovian age and E. holmi from the Late Wenlock age were described in 1934 by Leif Størmer. Additionally, he described four new English species, E. marstoni, E. spatulatus, E. megalodon and E. brodiei. E. marstoni (honoring Alfred Marston, responsible for several collections of fishes and eurypterids) is based on incomplete specimens, which together represent an almost complete chelicera. The holotype (BMNH 43790, in the British Museum of Natural History) consists of a free ramus, and the paratype (BMNH 43805, in the same museum as the holotype) includes most of the fixed ramus, that is very slender and tapering to the curved distal end. A large tooth is present in the midsection of the ramus. The end of the ramus is broken, but probably ended in a double tooth. This feature is also present in E. brodiei, only known from one specimen (FMNH 89411, located in the Field Museum of Natural History) which consists of an uncrushed chela. Its specific name honors Peter Bellinger Brodie, whose collections of eurypterids have helped the scientific community. E. spatulatus is remarkable by its shovel-or fan-shaped telson, which is wider than long and gives it the specific name. These telsons are rare, and have only been seen in E. grandis. E. serratus was designated a junior synonym of Pterygotus carmani in 2025. In 1971, E. serricaudatus and E. carinatus from the Early Wenlock age in Sweden were described by Kjellesvig-Waering. They were the only pterygotids in the area. The holotype of E. serricaudatus consists of the fixed ramus with acute termination of a large chelicera. This species is remarkable due to the group of diagonal opposing teeth of the ramus. This species differs from E. osiliensis (species in which E. serricaudatus was erroneously included) by its outwardly bowed rami, unlike the straight rami of E. serricaudatus. In addition, the shape and grouping of the teeth are completely different between them. The holotype of E. carinatus is the fragment of the base of a telson that measures in length and in width that indicates that the complete telson measured of estimated length, of maximum width. This species differs from others in having narrow lateral lobes, in having coarse striations on the cordated area and in its carinated (keel-like) telson, which is what gives the specific name. In 2025, E. carinatus was designated a junior synonym of E. serricaudatus. However, it has been questioned whether chelicerae serve as a factor for the distinction between genera since their morphology is dependent on lifestyle and vary throughout ontogeny (the formation and individual development of an organism), although they could be acceptable for the differentiation between species. Therefore, Truncatiramus was later recognized as representing a synonym of Erettopterus. In 2023, the new species Erettopterus qujingensis was described, from the upper Silurian (Pridolian) Yulongsi Formation of Yunnan Province, China. ==Classification==
Classification
Erettopterus is classified as part of the pterygotid family of eurypterids, and they were likely not capable of walking on land. What sets Erettopterus apart from the other pterygotids is the morphology of its bilobed telson, which is different from the telson of Ciurcopterus, with a dorsal median carinae, and that of Pterygotus, Jaekelopterus and Acutiramus, which form a short spine. ==Paleoecology==
Paleoecology
The chelicerae of the pterygotids were clearly adapted to be used for active prey capture and more similar to the claws of some modern crustaceans, with well developed teeth on the claws, than to the chelicerae of other eurypterid groups. Another feature distinguishing the group from other eurypterid groups were their flattened and expanded telsons, likely used as rudders when swimming. The cheliceral morphology and visual acuity of the pterygotid eurypterids separates them into distinct ecological groups. The primary method for determining visual acuity in arthropods is by determining the number of lenses in their compound eyes and the interommatidial angle (shortened as IOA and referring to the angle between the optical axes of the adjacent lenses). The IOA is especially important as it can be used to distinguish different ecological roles in arthropods, being low in modern active arthropod predators. The vision of Erettopterus was similar to that of the more basal pterygotoid Slimonia and more acute than the more derived Acutiramus though was not as acute as the vision of apex predators Jaekelopterus and Pterygotus or modern active predatory arthropods. Additionally, the large chelicerae of Erettopterus suggest that it was a generalized feeder and not a highly specialized predator and that it used its chelicerae (frontal appendages) to grasp. The teeth on the chelae fingers of Erettopterus were mostly short and suitable for holding prey, and could have effectively sliced up the prey. The deposits of the Rootsikula Formation in Saaremaa in which fossils of E. laticauda and E. osiliensis have been found shelter various faunas of eurypterids such as Mixopterus simonsi, Strobilopterus laticeps and Eysyslopterus patteni. Fossil remains of indeterminate osteostracids and thelodontids have also been found. Other fossils have also been found in different places, such as Great Britain, Canada, Scandinavia and the United States. ==See also==
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