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Tshekardocoleidae

Tshekardocoleidae is an extinct family of stem group beetles, known from the Early Permian. They represent some of the earliest known beetles. They first appeared during the Cisuralian, before becoming extinct at the beginning of the Guadalupian. A claimed Jurassic record is doubtful. Like other primitive beetles, they are thought to have been xylophagous. The oldest known beetle, Coleopsis, was originally assigned to this family, but is now assigned to its own family Coleopsidae.

Taxonomy
Kirejtshuk (2020) included the following genera in Tshekardocoleidae • †AvocoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Avocoleus fractus (Type species) • †Avocoleus neglegens • †BoscoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Boscoleus blandus (Type species) • †EocoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Eocoleus scaber (Type species) • †MoravocoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Moravocoleus permianus (Type species) • †PermocoleusMidco Member (Early Permian, Artinskian, ); Noble County, Oklahoma, USA • †Permocoleus wellingtoniensis (Type species) • †ProsperocoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Prosperocoleus prosperus (Type species) • †RetelytronObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Retelytron conopeum (Type species) • †SylvacoleodesTshekarda site (Early Permian, Kungurian, ); Suksun, Perm Krai, Russia • †Sylvacoleodes admirandus (Type species) • †SylvacoleusTshekarda site (Early Permian, Kungurian, ); Suksun, Perm Krai, Russia • †Sylvacoleus richteri (Type species) • †Sylvacoleus sharovi • †TshekardocoleusTshekarda site (Early Permian, Kungurian, ); Suksun, Perm Krai, Russia • †Tshekardocoleus magnus (Type species) • †Tshekardocoleus minor • †UmoricoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Umoricoleus perplex (Type species) • †VotocoleusObora site (Early Permian, Sakmarian, ); Moravia, Czech Republic • †Votocoleus submissus (Type species) The genus Uralocoleus had been placed in the family by some authors, but the type species Uralocoleus splendens was considered incertae sedis by Kirejtshuk (2020), as the type specimen is in bad condition and cannot be reliably compared with other members of the family, and the second species placed in the genus "Uralocoleus" ultimus was rejected from placement in Uralocoleus and listed as "genus incertus" ultimus , because it was based on an isolated elytron that could either belong to a beetle or a cockroach. The genus Dictyocoleus, described from the Jurassic and containing only the type species Dictyocoleus jurassicus , was originally placed in the family but was excluded by Kirejtshuk (2020). == References ==
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