Originally the reserve stood in a vast dryland steppe that stretched from the
Azov Sea and
Black Sea north for 500-700 km. Over times most of this area was converted to agriculture and only a few small sectors left in an uncultivated state. There are four main branches of the reserve, each protecting a different representative type of steppe for the region. • Khomutov Steppe. Located in
Kalmiuske Raion, near border with Russia. Established in 1921. Reportedly occupied by separatists in 2014. (1,030 hectares) • Stone Tombs •
Chalk Flora. Limestone cliffs along the right bank of the Donets River, supporting pine and oak forests. (1,134 hectares) • Kalmius Reserve. Rocky steppe (560 million year old granite) on floodplain of the
Kalmius River. (580 hectares) in the reserve ==Climate and ecoregion==