The Low-Vs served much of their time mainly on the
Lexington Avenue Express and
7th Avenue–
180th Street Bronx Park Express services, with a few additionally on the Broadway–7th Ave.–Van Cortlandt Park 242nd St. Express by interlining due to an insufficient number of High-Vs filling out the assigned services. At times, several appeared on the Queens Flushing and Astoria services as well. In 1952, with the shuffling of equipment to save on crews, the few Low-Vs on the Broadway Express were no longer needed for that service. A few, however, were used in Lenox Ave. Local service on weekends. In late 1958 through early 1959, several Low-Vs, following the Flivvers and regular Steinways, appeared on the two West Side Locals, Broadway and Lenox Ave., to replace the last of the High-Vs still in service. Around 1954–5, one car (5417) was fitted with additional light fixtures for brighter lighting, but the decision was made to instead rewire a few other cars with improved circuitry as with the IND R1–9s. Six cars were so equipped. These were 4800, 4953, 5095, 5285, 5416, 5440. Two cars (5032, 5130) had a lowered ceiling and hand stanchions in place of the usual straps. From August 1959 through December 1960, roughly 25 cars that had been modified for operation on the BMT Culver and Franklin Ave. Shuttle, by adding extensions on their sides at floor level to fill in the wider platform gaps on the
B Division; this lasted only for a year and a half, after which the cars were returned to the IRT. Most were placed in work service, surviving for many years, but 3 of these cars were actually returned to IRT passenger service and were scrapped in due course as with other Low-Vs at the time. After 47 years of service, most of the Low-V cars were retired in 1963, spending their last days on the Lexington–Jerome Ave. Express line. Some cars, however, remained in service on the 7th Avenue–Lenox Express until February 1964. There were persistent reports of those appearing in those services later that year. A very small number of Low-Vs used on the
Bowling Green–South Ferry shuttle that replaced the Deck Roof High-Vs - the previous fleet - in 1956 remained on the shuttle as late as August 1964. A handful number of trailer cars remained on the fragment of IRT Third Avenue Line in the Bronx to finish out their years until November 3, 1969. Many Low-Vs were converted into work cars following their retirement from revenue service. They were eventually replaced by retired SMEE cars converted into work cars and were eventually all scrapped (but see below). ==Preservation==