Valley elderberry longhorn beetles are stout-bodied. Males range in length from about 1.25–2.5 cm (½–1 in; measured from the front of the head to the end of the
abdomen) with
antennae about as long as their bodies. Females are slightly more robust than males, measuring about 1.9–2.5 cm (¾–1 in), with somewhat shorter antennae. The species is sexually dimorphic with adult males having red-orange
elytra (wing covers) with four elongate spots and margins. The red-orange fades to yellow on some museum specimens. As for the adult females, they have a dark-colored metallic green to black elytra with reddish margins. The four stages in the animal's life are:
egg,
larva,
pupa, and
adult. The species is nearly always found on or close to its host plant,
elderberry (
Sambucus species). Adults appear from the pith of elderberry at the start of the species' mating season in the spring, as the shrub begins to bloom. They feed on the leaf foliage, flowers, and nectar of the host plant before reproducing, and are active from March to June; this interval coincides with the blooming period of elderberry shrubs. In this period, the beetles mate and the females lay their eggs on the bark of living elderberry plants. The newly hatched larvae, or instar, burrow into the stems. The larval stage may last up to 1 to 2 years, in which they feed upon the pith as they grow. Before the pupal stage, the larvae bore exit holes through the surface, then sealing them with wood shavings. After which the larvae enter the pupal stage, returning to their chambers to undergo metamorphosis, later transforming into adults. The compound may be used as a lure for efficient monitoring of the valley elderberry longhorn beetle. ==Critical habitat==