Vulpes

Within Vulpes, 12 separate extant species and four fossil species are described: ==Early history==

The oldest known fossil species within Vulpes is V. riffautae, dating back to the late Miocene of Chad, which is within the Neogene. The deposits where these fossils are found are about 7 million years old, which might make them the earliest Canidae in the Old World. They are estimated to have weighed between 1.5 and 3.5 lb. V. skinneri, from the Malapa Fossil Site from South Africa, is younger than V. riffautae by roughly 5 million years, and shows up in the early Pleistocene. Two other extinct, less documented fossils are known: V. praeglacialis and V. hassani. V. praeglacialis was discovered in the Petralona Cave in Chalkidiki, Greece. The age of the deposits (Early Pleistocene) makes it the earliest occurrence of Vulpes in Europe. V. hassani is found in a Miocene-Pliocene deposit in northwestern Africa. which lends support that the close phylogenetic clustering of Rüppels and Red foxes is the result of recent introgressive hybridization rather than recent speciation. In the Pleistocene, Vulpes had a fairly wide distribution, with eight species found in North America. Of these eight, six are not fossil, and three species still remain in North America (V. velox, V. macrotis, and V. vulpes). The remaining three moved on to sections of Africa over time. V. stenognathus is extinct, but has extant sister taxa including V. chama, V. rueppellii, V. velox, and V. vulpes, which fits with these species all evolving together in North America. Fossil species • †Vulpes hassani • †Vulpes odessana • †Vulpes praeglacialis - Kormos (found in Petralona Cave, Greece) • †Vulpes qiuzhudingi (2014) • †Vulpes riffautae - Late Miocene • †Vulpes rooki • †Vulpes skinneri • †Vulpes stenognathus ==Description==

(Vulpes lagopus) in winter pelage. True foxes are small to medium-sized animals, usually smaller than other canines, such as wolves, dogs, and jackals. For example, the largest species, the red fox, weighs on average 4.1–8.7 kg and the smallest species, the fennec fox, weighs only 0.7–1.6 kg. They have long, dense fur, and a bushy, rounded tail that is at least half as long, or fully as long as, the head and body. They have a rather long body with shorter limbs, a long, narrow muzzle, and large, pointed ears. The forelimbs have five toes, while the hind legs have only four. Their skulls tend to be light, slender and elongated with a weak or non-existent sagittal crest. Vulpes species have vertically slit pupils, which generally appear elliptical in strong light like those of cats, which provide them with significant advantages. Like most canids, true foxes have a muscular body, powerful jaws, and teeth for grasping prey. Blunt claws are especially useful for gripping the ground while tracking down their prey. Some species have a pungent "foxy" odor, arising mainly from a gland located on the dorsal surface of the tail, not far from its base. The length, color and density of the fur of fox species differ. Fennec foxes (and other desert-adapted fox species such as Vulpes macrotis) have large ears and a short coat to keep the body cool. On the other hand, the Arctic fox has small ears and a thick, insulating coat to keep the body warm. A solid color coat is seen in most animals, but there are occasions where the coat color varies over the year to enhance camouflage against the current seasons landscape. The Arctic fox's tail-tip is of the same color as the rest of the tail (white or blue-gray). Blanford's fox usually possesses a black-tipped tail, but a small number of specimens (2% in Israel, 24% in the United Arab Emirates) possess a light-tipped tail. The other foxes in this group (Bengal, Cape, corsac, fennec, kit, pale, and swift) all possess black-tipped or dark-tipped tails. ==Distribution and habitat==

The range of the genus is very wide, present in a wide variety of habitats, from the desert to the Arctic, and from high altitudes in the mountains to open plains. True foxes are opportunistic and thrive anywhere they can find food and shelter. They are also widespread in suburban and urban areas, where they can take advantage of human food supplies; however, they prefer to stay away from large industrial areas. In certain areas, foxes tend to do better where humans are present, including in many agricultural landscapes, forests and patchy woodlands. ==Behavior and ecology==

Most true foxes are nocturnal, but they can be active during the morning and dusk and occasionally hunt and scavenge in daylight during winter. Many fox species are solitary or nomadic, living most of their lives on their own, except for the mating season, when they have a monogamous relationship with a partner. Some live in small family groups, others are more gregarious. Vulpes have a high variation in social organization between species and populations. Their hierarchical society usually depends on population densities. Cape foxes likely have a matriarchal social organization. Diet This genus is omnivorous and prone to scavenging. The foods of choice for Vulpes consist of invertebrates, a variety of small vertebrates, grasses, and some angiosperms. The typical intake per day is about 1 kg. True foxes exhibit hoarding behavior or caching where they store away food for another day out of sight from other animals. Predators Adult foxes have very few predators except coyotes, bears, and wolves, depending on the location. Juvenile foxes face a wider range of threats from small carnivores and large birds of prey, such as eagles. Breeding season varies between species and habitat, but they generally breed between late December and late March. Born deaf and blind, kits or cubs require their mother's milk and complete supervision for the first four to five weeks out of the womb, but begin to be progressively weaned after the first month. Once fully weaned, kits seek out various insects. The parents supplement this diet with a variety of mammals and birds. During early to middle July, the kits are able to hunt on their own and soon move away from their parents. ==Relationship with humans==

Domestication The silver fox is a melanistic form of the wild red fox. Though rare, domestication has been documented in silver foxes. The most notable experiment was conducted in Novosibirsk, Russia, at the Siberian Institute of Cytology and Genetics. In this study, generations of silver foxes were divided into those with friendly traits and those with unfriendly traits. After 50 years, the friendly foxes developed "dog-like" domesticated traits such as spots, tail wagging, enjoyment of human touch, and barking. Fox hunting Fox hunting was started in the United Kingdom in the 16th century that involves tracking, chasing, and killing a fox with the aid of foxhounds and horses. It has since then spread to Europe, the United States, and Australia. Vulpes in culture and literature ==References==