One of the earliest descriptions of the biodiversity in the Indo-Australian Archipelago dates back to 1521 when Venetian explorer
Pigafetta recorded the biological contrasts between the Philippines and the
Maluku Islands (Spice Islands) (on opposite sides of Wallace's Line) during the continuation of the voyage of
Ferdinand Magellan, after Magellan had been killed on
Mactan. Later on, the English navigator
G. W. Earl published his observations in faunal differences between the islands in the Indo-Australian archipelago. In 1845, G. W. Earl described how shallow seas connected islands on the west (
Sumatra,
Java, etc.) with the Asian continent and with similar wildlife, and islands on the east such as
New Guinea were connected to Australia and were characterized by the presence of
marsupials. These early investigations assisted Wallace in developing his theories about the biogeography which he stated publicly in his 1859 paper after extensively traveling the region. The proposal of the line, however, was not the main objective of Wallace's endeavours: His primary purpose was in fact to understand the geological phenomena and the colonization events that caused the boundaries in faunal distribution in the region through the development of his theories of evolution and biogeography. On the other hand, the lack of knowledge of
tectonic plates, and the uncertainty about biodiversity in the
Philippines, left Wallace with some contradicting points he had to deal with concerning his theory on biogeography. Huxley studied the distribution of
gallinaceous birds in the archipelago, and noticed that species in the Philippines were remarkably distinct from those in the Asiatic realm. Based on that, he redrew Wallace's boundary placing it to the west of the Philippines and named it "Wallace's Line", although Wallace himself had refused to include the Philippines on the east side of the line. •
Mayr's faunal balance (1944) In addition, several smaller transition sub-regional boundaries were also proposed. for instance, evaluated the distribution of land mammals, birds, and amphibians in Wallace's realms and concluded that the boundaries suggested by Wallace remain valid. Ali
et al. (2020), in a different attempt, studied the fauna of
Christmas Island and indicated that most of the ancestral colonizers of the island's land mammals and amphibians disappeared from the
Lombok Strait. Therefore, they propose a re-conformation of Wallace's Line so that Christmas Island would be sited on the Australasian side of the biogeographical divide, instead of the oriental side. ==Biogeography==