The fossil fauna and flora of the Norwich Crag have been studied since the early 19th century for
biostratigraphic and
palaeoclimatic interpretation. Taxa used include plants, molluscs,
foraminifers, mammals and
dinoflagellates. They provide evidence for a general climatic cooling trend from the
Pliocene into the
Pleistocene. The historic chronostratigraphic correlations and palaeoenvironmental interpretations based on biostratigraphy (local and continental) have been criticised as poorly defined and unreliable by Riches (2012). Vertebrate fossils tend to be concentrated in the basement bed Species of warm and cold substages are represented, sometimes mixed together; this may pose problems for palaeoecological interpretation.
Mammals Land mammal fauna From a biostratigraphic point of view, the terrestrial mammal fauna of the Norwich Crag belongs to the Late
Villafranchian European Land Mammal Age and is correlated with
European Neogene Mammal Zone MN17. The richest and best-preserved assemblage is found in deposits of Antian/Bramertonian age at Easton Wood,
Covehithe. Taxa collected from this site include the mammoth
Mammuthus meridionalis and the gomphothere
Anancus arvernensis, horse
Equus robustus, deer
Eucladoceros falconeri and
Eucladoceros sedgwicki, gazelle
Gazella anglica, beaver
Trogontherium sp., wolf
Canis etruscus, voles
Mimomys pliocaenicus,
Mimomys reidi and
Mimomys tigliensis, and lemming
Lemmus kowalskii. Bramerton has yielded
holotypes of the extinct otter
Enhydra reevei and the gazelle
Gazella daviesii. Vole fossils from the Norwich Crag contribute to the
‘vole clock’ used for biostratigraphic zonation of Pliocene and Pleistocene sediments.
Marine mammal fauna Marine taxa from Easton Wood include
Orca and the walrus
Alachatherium cretsii. The albatross
Phoebastria anglica has been recorded from Easton Wood, Covehithe.
Reptiles and Amphibians No
reptile and
amphibian fossils have yet been recorded from the Norwich Crag.
Fish Fossil marine fish from the Norwich Crag include genera
Chrysophrys sp. (a
snapper),
Acipenser (
sturgeon) and
Notidanus (a
cow shark), and species
Platax woodwardi (a batfish),
Gadus morhua (Atlantic cod),
Rhinoptera woodwardi (a cownose ray) and
Raja clavata (thornback ray). Teeth of the freshwater fish
Esox lucius (northern pike) have been found at Bramerton. However they show evidence of considerable transportation, and consequent mixing of faunal assemblages from the earlier Red Crag Formation. The use of molluscan fossils for climatic reconstruction has proved problematic. For instance species found together at several levels in the Ludham borehole notably
Serripes groenlandicus (Greenland cockle) and
Calyptraea chinensis (Chinese hat snail) have notably different climatic tolerances today (arctic and lusitanic respectively). Further work on Norwich Crag sediments at
Bramerton, Norfolk, allowed a
Bramertonian stage to be identified, characterised by temperate forest with
Quercus,
Carpinus and
Alnus. Correlation was made with the pollen assemblage from Chillesford. Specimens of fossil wood have occasionally been found, for example
pyritised pine and
oak at Holton.
Foraminifers Fossil
foraminifera provide important evidence for climatic and environmental interpretation and stratigraphic correlation in the Norwich Crag. The Ludham borehole has provided the most complete foraminiferal record, and has allowed the designation of seven biozones. Further biozones were identified at Bramerton and Easton Bavents, and assemblages identified at a number of other Norwich Crag sites, permitting correlation with the more complete sequence in the Netherlands. == Dating and correlations ==