Greater Anglia Production of the first unit for Greater Anglia began in February 2018 and was completed in September 2018. Early on, it had been intended for the type to enter service during 2019, but the fleet's introduction had to postponed into the following year due to software issues with the onboard Train Control Management System (TCMS). Testing of the first unit commenced on 17 March 2020, on the London Liverpool Street to Southend line. Both the testing and production programmes were impacted by the
COVID-19 pandemic that took hold during 2020. On 8 June 2020, operational use of the Class 720 was formally authorised by the
Office of Rail and Road (ORR). On 16 December 2020, they started service on the Braintree branch, and on 23 December 2020, the Class 720s were rolled out on Southminster services. In February 2021, it was reported that driver training had begun for their introduction to the Cambridge route. That same month, Ian McConnell, Greater Anglia's franchise and programmes director, noted that the Class 720 was already running relatively well and free of any meaningful technical issues, with challenges with the onboard software issues being the chief concern with the type's deployment at that point. On 25 August 2021, Class 720s entered service on the West Anglia mainline and Hertford East branch. On 13 September 2021, the 720s started services to Harwich Town and Walton-on-the-Naze. By April 2023, the fleet had replaced a wide number of trains used on suburban routes, with the final 720 being delivered in spring 2025.
c2c Class 720s for c2c were planned to enter service in 2021, On 6 October 2022, the
Southend Echo quoted a c2c spokesperson as saying "trains are currently undergoing rigorous testing. We will update our customers as soon as we have more information as to when they will be entering passenger service”. In March 2023, c2c's managing director confirmed in an interview that it was planned that the Class 720s would enter service later that year. On 2 June 2023, it was reported that the new trains would come into passenger service in September. The first c2c 720s duly entered service on 27 September 2023. The new trains have been praised by local commuters. ==Fleet details==