Caledonian Sleeper Virgin Trains contracted were replaced by
English, Welsh and Scottish Railway (EWS) Class 90s in March 1998. Their operation on the route ended in October 2019.
East Coast Main Line In the late 1990s, GNER hired Class 90s to stand in for Class 91s on London Kings Cross to Leeds services; 90024 was repainted into their dark blue livery. In 2016, DB Cargo Class 90s were returned to the East Coast Main Line on
Virgin Trains East Coast's London to , Leeds and services, whilst the Class 91 locomotives were overhauled. These continued to be operated by
London North Eastern Railway until 9 June 2019.
Greater Anglia As part of their
East Anglia franchise bid,
National Express East Anglia provided a fleet plan to replace the ageing Class 86 locomotives on the
Great Eastern Main Line with the more powerful and newer Class 90 locomotives, hoping this would improve performance and reduce operating costs. The Class 90s were to become available due to Virgin Trains' plan to replace all locomotive-hauled passenger trains with
Class 390 Pendolino units, freeing up their 15 Class 90 locomotives. 90001-015 were delivered progressively to
Crown Point TMD in
Norwich. These locomotives had their original louvered horn covers replaced with a single plate design with V-patterned holes, matching the
Mark 3 DVTs.
Abellio Greater Anglia took on the franchise in February 2012 and continued to operate the locomotives. In January 2020, the
Class 745 FLIRT sets began entering service to replace the Class 90 sets. Following the introduction of these units, the locomotive-hauled sets were withdrawn from service, with the last set running its last services on 24 March 2020. Thirteen moved to Freightliner to replace the Class 86s, while the other two were purchased by Locomotive Services Limited. The first locomotives to be replaced were the elderly Class 86/2s and some of the less reliable Class 87 locomotives. However, since the Class 90 fleet was relatively small and subsequently non-standard, it was decided to retain the larger Class 87 fleet in preference to the Class 90s. Therefore, from March 2004, Virgin started to return its Class 90 fleet to the ROSCO, allowing the locos to transfer to the new
One (later National Express East Anglia) operator. Following the
Grayrigg derailment and subsequent scrapping of 390033 in 2007, Virgin Trains had the need for an additional set as a stop gap before another Pendolino set was built. As a result, Virgin used Class 90 locomotives hired from DB Schenker, and later Freightliner, along with a rake of
Mark 3 coaching stock and a DVT. Nicknamed the
Pretendolino, this received reupholstered seating, power points,
wi-fi and a full external re-paint at
Wabtec,
Doncaster, in 2009. Virgin used this set, with a Class 90 locomotive hired from Freightliner, on a Friday relief service between Euston and Crewe, via Birmingham, until December 2012. It was hired out as a charter train and was used sometimes on London-Birmingham services in the event of a Pendolino shortage. From the December 2013 timetable change, Virgin Trains used the Mark 3 set once again on London-Birmingham services on a Thursday and Friday only basis; Class 90s were leased from
Direct Rail Services, which were sub-leased from DB Schenker. ==Fleet==