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Eusarcana

Eusarcana is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Eusarcana have been discovered in deposits ranging in age from the Early Silurian to the Early Devonian. Classified as part of the family Carcinosomatidae, the genus contains three species, E. acrocephalus from Scotland and E. scorpionis and E. spiniferum from the United States.

Description
Eusarcana can be differentiated from other eurypterids by the considerably narrow border between the prosoma (head) and opisthosoma (abdomen), which is particularly thin considering the subsequent broad and large ellipse-shape of the abdomen. The postabdomen (or tail) also narrows rather quickly from the preceding segments. Further features distinctive of the genus include that the carapace (segment covering the head) is clearly triangular in shape, with eyes placed on the rim of it and positioned forwards, the fact that all walking legs possess spines and that they decrease in length the further back they were placed as well as the cylindrically shaped and sharp telson (the posteriormost division of the body). In comparison with many other eurypterids, Eusarcana was a rather large animal, The appearance of Eusarcana is somewhat odd in comparison with other eurypterids, not only in its overall shape and proportions but also in that the surface of its exoskeleton is covered in small scale-like ornamentation that is circular in shape, crowded and small in size which differentiates it from other eurypterids in which such ornamentation is usually triangular. == History of research ==
History of research
Original description and subsequent discoveries Eusarcana was first described as "Eusarcus" by the American geologists August R. Grote and William Henry Pitt based on fossils recovered from the Pridoli-age Buffalo Waterlime of New York State. This name derives from the Ancient Greek εὖ, (eu-) meaning "true", and σάρξ (sarx), meaning "flesh", meaning "true flesh". The designated type species was E. scorpionis. Though Grote and Pitt did not provide a generic diagnosis for the genus, the species was well diagnosed with a number of distinctive characters. Furthermore, the genus of Grote and Pitt was seemingly based solely on outlines and shape, which prompted some researchers, such as the prominent English geologist Henry Woodward, to regard the genus as lacking generic characters and as such being invalid, referring E. scorpionis to Eurypterus on the grounds that several British species of Eurypterus, notably E. scorpiodes and E. punctatus (today recognized as species of Carcinosoma), were similar in shape. In 2014, American paleontologist James Lamsdell suggested that E. obesus may represent a juvenile of the related and contemporary Carcinosoma scorpioides. Unlike other species of Eusarcana, E. obesus does not appear to possess spines on its appendages. Carcinosoma scorpioides was referred to the new genus Cruinnopterus by Lamsdell (2025), and E. obesus was designated as a junior synonym of that species.'''''' Lamsdell (2025) also transferred the species Carcinosoma spiniferum to Eusarcana.'''' E. spiniferum is known from the Silurian of New York, from a single prosomal appendage that preserves some elongated spines. The appendage is much less robust than the appendage of Carcinosoma newlini, the Carcinosoma type species, and is more similar to that of E. scorpionis.'''' Recognition as preoccupied name It was first in 1934, 59 years after its original description, that Eusarcus was recognized as a name preoccupied by a harvestman. The Norwegian geologist Leif Størmer proposed that the name of the taxon should be next oldest available and valid name for the genus, Carcinosoma. During the preparation for his paper on the issue, Størmer also discussed the situation with fellow Norwegian researcher Embrik Strand, who helped confirm that Carcinosoma was not preoccupied. Strand would subsequently propose the replacement name Eusarcana in 1942, despite the problem having been dealt with by Størmer, who he had been in contact with eight years earlier. The reasons for proposing the name during the circumstances of the time remains unknown, but contemporary researchers critiquing Strand for his studies in systematics and an apparent desire to name as many taxa as possible may explain the situation somewhat. As it was seen as completely unnecessary at the time, Strand's Eusarcana was overlooked and not even mentioned in subsequent eurypterid studies. The naming of Eusarcana was one of many contributions to nomenclature by Strand seen as unhelpful today. His journal Folia Zoologica et Hydrobiologica would later re-emerge within the field to create all kinds of systematic problems that could have been avoided. Strand was also notorious for applying new species names to incomplete or poorly preserved fossils. In 1964, American paleontologists Kenneth Edward Caster and Erik N. Kjellesvig-Waering recognized Eusarcus and Carcinosoma to be distinct genera when revising the superfamily Carcinosomatoidea, and coined the replacement name Paracarcinosoma to designate the species previously assigned to Eusarcus. E. scorpionis was designated the type species. Caster and Kjellesvig-Waering made no mention of Embrik Strand or Eusarcana, and they were likely not aware of the existence of the previous name. With Eusarcana all but forgotten, all subsequent researchers used Paracarcinosoma for the genus. In 2012, American paleontologists Jason A. Dunlop and James Lamsdell noted that whilst the name Eusarcana had been completely unnecessary (and of questionable validity, as Strand did not designate a type species) at the time of its creation, it is the oldest available valid name for the taxon and as such should constitute its name under the rules of priority, despite Paracarcinosoma being more widely used. As such, Paracarcinosoma was designated as a junior synonym, with all three species assigned to it being transferred to Eusarcana. == Classification ==
Classification
'', a related carcinosomatid species Eusarcana is classified as part of the family Carcinosomatidae, a family within the superfamily Carcinosomatoidea, alongside the genera Carcinosoma, Eocarcinosoma, Rhinocarcinosoma and possibly Holmipterus. The cladogram below is adapted from a larger cladogram (simplified to only display the Carcinosomatoidea) in a 2007 study by eurypterid researcher O. Erik Tetlie, in turn based on results from various phylogenetic analyses on eurypterids conducted between 2004 and 2007. The second cladogram below is simplified from a study by Lamsdell et al. (2015). Tetlie (2007) 'Lamsdell et al. (2015)' {{clade| == Paleobiology ==
Paleobiology
The features of Eusarcana indicate that its lifestyle was completely different from many other swimming eurypterids, such as Eurypterus. When walking, Eusarcana would likely have lifted the eye-bearing frontal part of its head above the ground, as indicated by the larger size of the forward walking legs and the placement of the eyes. In contrast, Eurypterus would have kept its head, shovel-shaped and broad, down to the ground while walking. Furthermore, the walking legs of Eusarcana are more powerful than those of Eurypterus in general. The legs decrease in strength the further back they are, indicating that there was an emphasis on lifting the front of the carapace. In Eurypterus the legs are the longest the further back they are, which indicates that it would have had an emphasis on keeping the head down. Complex eurypterid faunas, compromising several different species in different ecological roles, are typical of the period. The Silurian-aged fauna of the same general area, also preserving E. acrocephalus, were home to other eurypterids Erettopterus, Pterygotus and Acutiramus, a less diverse assemblage of trilobites (Otarion, Scharyia and Prionopeltis), crustacean Ceratiocaris, cephalopods (including Cycloceras and Corbuloceras), ostracods, gastropods, crinoids, conodonts and bivalves. In Silurian deposits of New York, E. scorpionis and E. spiniferium occur together with a diverse fauna of eurypterids composed of Buffalopterus, Dolichopterus, Erettopterus, Eurypterus, Pterygotus and Acutiramus. Also present were xiphosurans, crustaceans (Gonatocaris), ostracods, bivalves, gastropods and cephalopods. Carcinosomatid eurypterids such as Eusarcana were among the most marine eurypterids, known from deposits that were once reefs, some in lagoonal settings, and deeper waters. ==See also==
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