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Charles Scott Sherrington

Sir Charles Scott Sherrington was a British neurophysiologist. His experimental research established many aspects of contemporary neuroscience, including the concept of the spinal reflex as a system involving connected neurons, and the ways in which signal transmission between neurons can be potentiated or depotentiated. Sherrington himself coined the word "synapse" to define the connection between two neurons. His book The Integrative Action of the Nervous System (1906) is a synthesis of this work, in recognition of which he was awarded the Nobel Prize in Physiology or Medicine in 1932.

Biography
Early years and education Official biographies claim Charles Scott Sherrington was born in Islington, London, England, on 27 November 1857, and that he was the son of James Norton Sherrington, a country doctor, and his wife, Anne Thurtell. However James Norton Sherrington was an ironmonger and artist's colourman in Great Yarmouth, not a doctor, and died in Yarmouth in 1848, nearly 9 years before Charles was born. In the 1861 census, Charles is recorded as Charles Scott (boarder, 4, born India) with Anne Sherrington (widow) as the head and Caleb Rose (visitor, married, surgeon). He was brought up in this household with Caleb recorded as head in 1871, although Anne and Caleb did not marry until after the death of his wife in 1880. The relationship between Charles and his childhood family is unknown. During the 1860s the whole family moved to Anglesea Road, Ipswich, reputedly because London exacerbated Caleb Rose's tendency to asthma. Sherrington's origins have been discussed in several published sources: Chris Moss and Susan Hunter, in the Journal of Medical Biography of January 2018, presented an article discussing the potential origins of Charles Sherrington, i.e. whether he was born in India of unknown parents, or was the illegitimate child of Caleb Rose and Anne Sherrington. Erling Norrby, PhD, in Nobel Prizes and Notable Discoveries (2016) observed: "His family origin apparently is not properly given in his official biography. Considering that motherhood is a matter of fact and fatherhood a matter of opinion, it can be noted that his father was not James Norton Sherrington, from whom his family name was derived. Charles was born 9 years after the death of his presumed father. Instead Charles and his two brothers were the illegitimate sons of Caleb Rose, a highly regarded Ipswich surgeon." In Ipswich Town: A History, Susan Gardiner writes: "George and William Sherrington, along with their older brother, Charles, were almost certainly the illegitimate sons of Anne Brookes, née Thurtell and Caleb Rose, a leading surgeon from Ipswich, with whom she was living in College Road, Islington at the time that all three boys were born. No father was named in the baptism register of St James' Church, Clerkenwell, and there is no official record of the registration of any of their births. It was claimed they were the sons of a country doctor, James Norton Sherrington. However, it was with Caleb Rose that Anne and the three Sherrington boys moved to Anglesea Road, Ipswich in 1860 and the couple were married in 1880 after Caleb's first wife had died." Judith Swazey, in ''Reflexes and Motor Integration: Sherrington's Concept of Integrative Action'' (1969), quotes Charles Scott Sherrington's son, Carr Sherrington: "James N. Sherrington was always called Mr. and I have no knowledge that he was a Dr. either in law or in medicine... [He] was mainly interested in art and was a personal friend of J. B. Crone and other painters." Caleb Rose was noteworthy as both a classical scholar and an archaeologist. At the family's Edgehill House in Ipswich one could find a fine selection of paintings, books, and geological specimens. Through Rose's interest in the Norwich School of Painters, Sherrington gained a love of art. Intellectuals frequented the house regularly. It was this environment that fostered Sherrington's academic sense of wonder. Even before matriculation, the young Sherrington had read Johannes Müller's Elements of Physiology. The book was given to him by Caleb Rose. Sherrington entered Ipswich School in 1871. Sherrington played football for his grammar school, and for Ipswich Town Football Club; he played rugby for St. Thomas's, was on the rowing team at Oxford. During June 1875, Sherrington passed his preliminary examination in general education at the Royal College of Surgeons of England (RCS). This preliminary exam was required for Fellowship, and also exempted him from a similar exam for the Membership. In April 1878, he passed his Primary Examination for the Membership of the RCS, and twelve months later the Primary for Fellowship. In October 1879, Sherrington entered Cambridge as a non-collegiate student. The following year he entered Gonville and Caius College. Sherrington was a first-rate student. In June 1881, he took Part I in the Natural Sciences Tripos (NST) and was awarded a Starred first in physiology; there were nine candidates in all (eight men, one woman), of whom five gained First-class honours (Firsts); in June 1883, in Part II of the NST, he also gained a First, alongside William Bateson. Walter Holbrook Gaskell, one of Sherrington's tutors, informed him in November 1881 that he had earned the highest marks for his year in botany, human anatomy, and physiology; second in zoology; and highest overall. There, Sherrington worked on segmental distribution of the spinal dorsal and ventral roots, he mapped the sensory dermatomes, and in 1892 discovered that muscle spindles initiated the stretch reflex. The institute allowed Sherrington to study many animals, both small and large. The Brown Institute had enough space to work with large primates such as apes. Liverpool Sherrington's first job of full-professorship came with his appointment as Holt Professor of Physiology at Liverpool in 1895, succeeding Francis Gotch. Sherrington also influenced American pioneer brain surgeon Harvey Williams Cushing. Sherrington's philosophy as a teacher can be seen in his response to the question of what was the real function of Oxford University in the world. Sherrington said: "After some hundreds of years of experience we think that we have learned here in Oxford how to teach what is known. But now with the undeniable upsurge of scientific research, we cannot continue to rely on the mere fact that we have learned how to teach what is known. We must learn to teach the best attitude to what is not yet known. This also may take centuries to acquire but we cannot escape this new challenge, nor do we want to." Sherrington's teachings at Oxford were interrupted by World War I. When the war started, it left his classes with only nine students. During the war, he laboured at a shell factory to support the war and to study fatigue in general, but specifically industrial fatigue. His weekday work hours were from 7:30am to 8:30pm; and from 7:30am to 6:00pm on the weekends. Retirement Charles Sherrington retired from Oxford in the year of 1936. Sherrington's mental faculties were crystal clear up to the time of his sudden death, which was caused by a sudden heart failure at age 94. His bodily health, however, did suffer in old age. Arthritis was a major burden. Speaking of his condition, Sherrington said "old age isn't pleasant[,] one can't do things for oneself." The arthritis put Sherrington in a nursing home in the year before his death, in 1951. ==Family==
Family
On 27 August 1891, Sherrington married Ethel Mary Wright (died 1933), daughter of John Ely Wright of Preston Manor, Suffolk, England. They had one child, a son named Charles ("Carr") E.R. Sherrington, who was born in 1897. On weekends during the Oxford years the couple would frequently host a large group of friends and acquaintances at their house for an enjoyable afternoon. ==Publications==
Publications
The Integrative Action of the Nervous System Published in 1906, The book discussed neuron theory, the "synapse" (a term he had introduced in 1897, the word itself suggested by classicist A. W. Verrall), communication between neurons, and a mechanism for the reflex-arc function. In the original edition, each of the twelve chapters begins with one of the twelve zodiac signs; Sherrington discusses astrology in Fernel's time in Chapter 2. In his ideas on mind and cognition, Sherrington introduced the idea that neurons work as groups in a "million-fold democracy" to produce outcomes rather than with central control. ==Honours and awards==
Honours and awards
Sherrington was elected a Fellow of the Royal Society (FRS) in 1893. • 1922 President of the British Association for the year 1922–1923 • 1924 Order of Merit • 1927 Copley Medal • 1932 Nobel Prize for Physiology or Medicine At the time of his death Sherrington received honoris causa Doctors from twenty-two universities: Oxford, Paris, Manchester, Strasbourg, Louvain, Uppsala, Lyon, Budapest, Athens, London, Toronto, Harvard, Dublin, Edinburgh, Montreal, Liverpool, Brussels, Sheffield, Bern, Birmingham, Glasgow, and the University of Wales. Eponyms ;Liddell-Sherrington reflex: Associated with Edward George Tandy Liddell and Charles Scott Sherrington, the Liddell-Sherrington reflex is the tonic contraction of muscle in response to its being stretched. When a muscle lengthens beyond a certain point, the myotatic reflex causes it to tighten and attempt to shorten. This can be felt as tension during stretching exercises. ;Schiff-Sherrington reflex: Associated with Moritz Schiff and Charles Scott Sherrington, describes a grave sign in animals: rigid extension of the forelimbs after damage to the spine. It may be accompanied by paradoxical respiration – the intercostal muscles are paralysed and the chest is drawn passively in and out by the diaphragm. ;Sherrington's First Law: Every posterior spinal nerve root supplies a particular area of the skin, with a certain overlap of adjacent dermatomes. ;Sherrington's Second Law: The law of reciprocal innervation. When contraction of a muscle is stimulated, there is a simultaneous inhibition of its antagonist. It is essential for coordinated movement. ;Vulpian-Heidenhain-Sherrington phenomenon: Associated with Rudolf Peter Heinrich Heidenhain, Edmé Félix Alfred Vulpian, and Charles Scott Sherrington. Describes the slow contraction of denervated skeletal muscle by stimulating autonomic cholinergic fibres innervating its blood vessels. ==See also==
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