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Chenopodium

Chenopodium is a genus of numerous species of perennial or annual herbaceous flowering plants known as the goosefoot, which occur almost anywhere in the world. It is placed in the family Amaranthaceae in the APG II system; older classification systems, notably the widely used Cronquist system, separate it and its relatives as Chenopodiaceae, but this leaves the rest of the Amaranthaceae polyphyletic. However, among the Amaranthaceae, the genus Chenopodium is the namesake member of the subfamily Chenopodioideae.

Description
The species of Chenopodium (s.str., description according to Fuentes et al. 2012) They generally rely on alkaline soil. They are nonaromatic, but sometimes fetid (foul-smelling). The young stems and leaves are often densely covered by vesicular globose hairs, thus looking farinose. Characteristically, these trichomes persist, collapsing later and becoming cup-shaped. The branched stems grow erect, ascending, prostrate or scrambling. Lateral branches are alternate (the lowermost ones can be nearly opposite). The alternate or opposite leaves are petiolate. Their thin or slightly fleshy leaf blade is linear, rhombic or triangular-hastate, with entire or dentate or lobed margins. Inflorescences are standing terminal and lateral. They consist of spicately or paniculately arranged glomerules of flowers. Plants are monoecious (rarely dioecious). In monoecious plants flowers are dimorphic or pistillate. Flowers consist of (4–) 5 perianth segments connate, basally (at the base) or close to the middle, usually membranous margined and with a roundish to keeled back; almost always 5 stamens, and one ovary with 2 stigmas. In fruit, perianth segments become sometimes coloured, but mostly keep unchanged, somewhat closing over or spreading from the fruit. The pericarp is membranous or sometimes succulent, adherent to or loosely covering the seed. The horizontally oriented seeds are depressed-globular to lenticular, with rounded to subacute margin. The black seed coat is almost smooth to finely striate, rugulose or pitted. == Taxonomy ==
Taxonomy
The genus Chenopodium was described by Carl Linnaeus in 1753 (In: Species Plantarum, Vol. 1, p. 218–222). Type species is Chenopodium album. This generic name is derived from the particular shape of the leaf, which is similar to a goose's foot: from Greek χήν (chen), "goose" and πούς (pous), "foot" or (podion), "little foot". In its traditional circumscription, Chenopodium comprised about 170 species. • Chenopodium candolleanum (Syn.: Rhagodia candolleana) • Chenopodium carnosulumChenopodium chaldoranicumChenopodium cordobenseChenopodium cornutumChenopodium × covilleiChenopodium crusoeanumChenopodium cuneifoliumChenopodium curvispicatumChenopodium cyanifoliumChenopodium cycloides – sandhill goosefoot • Chenopodium × dadakovaeChenopodium desertorum – desert goosefoot • Chenopodium desiccatum – narrowleaf goosefoot • Chenopodium detestans – New Zealand fish-guts plant • Chenopodium diversifoliumChenopodium drummondii (Syn.: Rhagodia drummondii) • Chenopodium eastwoodiae – Eastwood's goosefoot • Chenopodium eremaeum (Syn.: Rhagodia eremaea) • Chenopodium erosum R.Br. • Chenopodium eustriatumChenopodium × fallaxChenopodium ficifoliiformeChenopodium ficifolium – fig-leaved goosefoot, small goosefoot • Chenopodium flabellifolium – San Martin Island goosefoot, flabelliform goosefoot • Chenopodium foggii – Fogg's goosefoot • Chenopodium fremontii – Fremont's goosefoot • Chenopodium frutescensChenopodium × fursajeviiChenopodium gaudichaudianumChenopodium giganteum D.Don (synonym Chenopodium formosanum ) – tree spinach, red quinoa, djulis • Chenopodium griseochlorinumChenopodium gruboviiChenopodium × gruelliiChenopodium haraeChenopodium hastatifoliumChenopodium × haywardiaeChenopodium hederiformeChenopodium hiansChenopodium hircinumChenopodium hoggarenseChenopodium howellii – Howell's goosefoot • Chenopodium hubbardiiChenopodium × humiliformeChenopodium iljiniiChenopodium incanum – mealy goosefoot • Chenopodium incognitumChenopodium indicumChenopodium iranicumChenopodium × jedlickaeChenopodium × jehlikiiChenopodium karoiChenopodium khorasanicaChenopodium lenticulareChenopodium × leptophylliformeChenopodium leptophyllum – narrowleaf goosefoot • Chenopodium × lincienseChenopodium lineatum – Mono goosefoot • Chenopodium littoreum – coastal goosefoot • Chenopodium lobodontumChenopodium loureiroiChenopodium luteorubrumChenopodium luteum – yellow goosefoot • Chenopodium × mendeliiChenopodium moquinianumChenopodium mucronatumChenopodium neomexicanum – New Mexico goosefoot • Chenopodium nesodendronChenopodium nevadense – Nevada goosefoot • Chenopodium nitensChenopodium nitrariaceum (F.Muell.) F.Muell. ex Benth. – nitre goosefoot • Chenopodium nitens – shiny goosefoot • Chenopodium novopokrovskyanumChenopodium nutans (Syn.: Rhagodia nutans) • Chenopodium oahuenseĀheahea (Hawaii) • Chenopodium obscurumChenopodium opulifolium Schrad. ex W.D.J.Koch & Ziz – grey goosefoot • Chenopodium pallescens – pallid goosefoot • Chenopodium pallidicaulekañiwa, "cañahua" • Chenopodium palmeri – Palmer's goosefoot • Chenopodium pamiricumChenopodium parabolicum (Syn.: Rhagodia parabolica) • Chenopodium × paradoxumChenopodium parryi – Parry's goosefoot • Chenopodium perttiiChenopodium petiolareChenopodium philippianumChenopodium phillipsianumChenopodium pilcomayenseChenopodium × podperaeChenopodium polygonoidesChenopodium × praeacutumChenopodium pratericola Rydb. – pale goosefoot, desert goosefoot, narrowleaf goosefoot • Chenopodium preissii (Syn. Rhagodia preissii) • Chenopodium × preissmanniiChenopodium × pseudoleptophyllumChenopodium × pseudostriatumChenopodium pueblenseChenopodium quinoa – quinoa • Chenopodium × reynieriChenopodium robertianum (Syn.: Rhagodia hastata) • Chenopodium ruiz-lealiiChenopodium sanctae-claraeChenopodium sancti-ambrosiiChenopodium sandersii – Sander's goosefoot • Chenopodium santosheiChenopodium scabricauleChenopodium simpsonii – Simpson's goosefoot • Chenopodium × smardaeChenopodium sonorense – Sonoran goosefoot • Chenopodium sosnowskyiChenopodium spegazziniiChenopodium spinescens (Syn. Rhagodia spinescens) • Chenopodium standleyanum – Standley's goosefoot • Chenopodium stenophyllumChenopodium striatiformeChenopodium subficifoliumChenopodium subglabrum – smooth arid goosefoot, smooth goosefoot • Chenopodium suecicum – green goosefoot • Chenopodium × thellungiiChenopodium tonkinenseChenopodium triandrum (Syn.: Rhagodia triandra) • Chenopodium × tridentinumChenopodium × trigonocarpumChenopodium trigonon (Syn.: Einadia trigonos) • Chenopodium twisselmannii – Twisselmann's goosefoot, high meadow goosefoot • Chenopodium ulbrichiiChenopodium ulicinumChenopodium × unariiChenopodium × variabile (C. album × C. berlandieri) • Chenopodium vulvaria – stinking goosefoot, notchweed • Chenopodium wahlii – Wahl's goosefoot • Chenopodium watsonii – Watson's goosefoot • Chenopodium wilsonii (Syn.: Rhagodia crassifolia) • Chenopodium × zahniiChenopodium zeroviiChenopodium zoellneri Excluded speciesBlitum (12 species): • Blitum bonus-henricus – Good King Henry, perennial goosefoot, poor-man's asparagus, Lincolnshire spinach, markery • Blitum californicum – California goosefoot, Indian lettuce • Blitum capitatum – strawberry blite, blite goosefoot, strawberry goosefoot, strawberry spinach, Indian paint, Indian ink • Blitum virgatum (Syn. Chenopodium foliosum) – leafy goosefoot • Chenopodiastrum (5 species): • Chenopodiastrum murale – nettle-leaved goosefoot • Chenopodiastrum simplex – giant seed goosefoot • Dysphania (about 43 glandular species, as C. botrys, C. carinatum, C. cristatum, C. melanocarpum, C. multifidium, C. pumilio and more) • Lipandra (one species): • Lipandra polysperma – many-seeded goosefoot • Oxybasis (5 species): • Oxybasis chenopodioides – small red goosefoot, saltmarsh goosefoot • Oxybasis glauca – oak-leaved goosefoot • Oxybasis rubra – red goosefoot, coastblite goosefoot • Oxybasis urbica – upright goosefoot • Teloxys (one species): • Teloxys aristataSuaeda australis – austral seablite (as C. australe, C. insulare) ==Ecology==
Ecology
Certain species grow in large thickets, providing cover for small animals. Goosefoot foliage is used as food by the caterpillars of certain Lepidoptera. The seeds are eaten by many birds, such as the yellowhammer (Emberiza citrinella) of Europe or the white-winged fairy-wren (Malurus leucopterus) of Australia. Goosefoot pathogens include the positive-sense ssRNA virusesapple stem grooving virus, sowbane mosaic virus and tobacco necrosis virus. ==Uses==
Uses
(C. quinoa) seeds The genus Chenopodium contains several plants of minor to moderate importance as food crops as leaf vegetables – used like the closely related spinach (Spinacia oleracea) and similar plants called quelite in Mexicoand pseudocereals. These include white goosefoot (C. album), kañiwa (C. pallidicaule) and quinoa (C. quinoa). On the Greek island of Crete, tender shoots and leaves of a species called krouvida () or psarovlito () are eaten by the locals, boiled or steamed. As studied by Bruce D. Smith, Kristen Gremillion and others, goosefoots have a history of culinary use dating back to 4000 BC or earlier, when pitseed goosefoot (C. berlandieri) was a staple crop in the Native American Eastern Agricultural Complex, and when white goosefoot was apparently used by the Ertebølle culture of Europe. Members of the eastern European Yamnaya culture also harvested white goosefoot as an apparent cereal substitute to round out an otherwise mostly meat and dairy diet c. 3500–2500 BC. There is increased interest in particular in goosefoot seeds today, which are suitable as part of a gluten-free diet. Quinoa oil, extracted from the seeds of C. quinoa, has similar properties, but is superior in quality, to corn oil. Oil of chenopodium is extracted from the seeds of epazote, which is not in this genus anymore. Shagreen leather was produced in the past using the small, hard goosefoot seeds. C. album was one of the main model organisms for the molecular biological study of chlorophyllase. Goosefoot pollen, in particular of the widespread and usually abundant C. album, is an allergen to many people and a common cause of hay fever. The same species, as well as some others, have seeds which are able to persist for years in the soil seed bank. Many goosefoot species are thus significant weeds, and some have become invasive species. We have recently gathered an abundant harvest of leaves from two or three plants growing in our garden. These leaves were put into boiling water to blanch them, and they were then cooked as an ordinary dish of spinach, with this difference in favour of the new plant, that there was no occasion to take away the threads which are so disagreeable in chicory, sorrel, and ordinary spinach. We partook of this dish with relish—the flavour—analogous to spinach, had something in it more refined, less grassy in taste. The cultivation is easy: sow the seed in April (October) in a well-manured bed, for the plant is greedy; water it. The leaves may be gathered from the time the plant attains 50 centimetres (say 20 inches) in height. They grow up again quickly. In less than eight days afterwards another gathering may take place, and so on to the end of the year. Safety Sphaeraphides occur in the leaves, stem, pith and mesophloem. ==Fossil record==
Fossil record
Chenopodium wetzleri fossil seeds of the Chattian stage, Oligocene, are known from the Oberleichtersbach Formation in the Rhön Mountains, central Germany. ==References==
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