It is speculated by paleontologists that during the
Eocene period hooved marsh dwellers carried their body weight mainly on two of the middle toes, which grew to equal size, becoming the
Artiodactyla or even-toed hooved animals. Before the end of the Eocene period the side toes of some had dwindled and practically disappeared (mainly in the form of a
dewclaw) while the basal pieces or
metapodium of the pair of supporting toes became fused together, thus producing the appearance of a cloven hoof. The mammal with a cloven hoof is an even-toed
ungulate of order Artiodactyla as opposed to the
odd-toed ungulates of Perissidactyla, such as the
horse, which have one toe, or the
rhinoceros, which has three toes. The five-toed ancestors of the earliest
Eocene had already developed feet that suggest odd-toed and even-toed descendants to the modern viewer. Even
Phenacodus, the most generalized of the early mammals, has a foot in which the central toe is somewhat larger than the others and could be placed in the division of odd-toed ungulates, Perissidactyla. ==Jewish dietary laws==