•
Copris acutidens Motschulsky, 1860 •
Copris afgoi Cambefort, 1992 •
Copris agnus Sharp, 1875 •
Copris algol Nguyen-phung, 1989 •
Copris amabilis Kolbe, 1914 •
Copris amyntor Klug, 1855 •
Copris anceus (Olivier, 1789) •
Copris andrewesi Waterhouse, 1891 •
Copris angusticornis Arrow, 1933 •
Copris angustus Nguyen-phung & Cambefort, 1986 •
Copris anomiopseoides Boucomont, 1924 •
Copris antares Ferreira, 1958 •
Copris arcturus Gillet, 1907 •
Copris ares Zelenka, 1993 •
Copris arimotoi Ochi & Araya, 1992 •
Copris arizonensis Schaeffer, 1906 •
Copris armatus Harold, 1869 •
Copris armeniacus Faldermann, 1835 •
Copris armiger Gillet, 1910 •
Copris arrowi Felsche, 1910 •
Copris aspericollis Gillet, 1910 •
Copris atropolitus Gillet, 1910 •
Copris bacchus Thunberg, 1787 •
Copris barclayi Ochi, Kon & Bai, 2009 •
Copris basilewskyi Ferreira, 1962 •
Copris basipunctatus Balthasar, 1942 •
Copris bellator Chevrolat, 1844 •
Copris bengalensis Gillet, 1911 •
Copris bidens Kolbe, 1893 •
Copris binodis •
Copris bootes Klug, 1855 •
Copris boucardi Harold, 1869 •
Copris bovinus Gillet, 1908 •
Copris brachypterus Nomura, 1964 •
Copris brittoni Ferreira, 1962 •
Copris caelatus (Fabricius, 1794) •
Copris caliginosus Kohlmann, Cano & Delgado, 2003 •
Copris cambeforti Nguyen-phung, 1988 •
Copris cambodiensis Ochi, Kon & Kawahara, 2008 •
Copris camerunus Felsche, 1904 •
Copris caobangensis Bui, Dumack & Bonkowski, 2018 •
Copris capensis Waterhouse, 1891 •
Copris cariniceps Felsche, 1910 •
Copris carinicus Gillet, 1910 •
Copris carmelita Fabricius, 1801 •
Copris cassius Péringuey, 1901 •
Copris celebensis Lansberge, 1886 •
Copris cheni Ochi, Kon & Bai, 2007 •
Copris chimalapensis Mora-Aguilar & Delgado, 2015 •
Copris ciliatus •
Copris colmanti Gillet, 1908 •
Copris complexus Nguyen-phung, 1989 •
Copris compressipennis Gillet, 1910 •
Copris confucius Harold, 1877 •
Copris coriarius Gillet, 1907 •
Copris cornifrons Boheman, 1860 •
Copris corniger Sahlberg, 1823 •
Copris corpulentus Gillet, 1910 •
Copris costaricensis Gahan, 1894 •
Copris costariensis Matthews •
Copris crassus Deschodt & Davis, 2015 •
Copris cribratus Gillet, 1927 •
Copris cribricollis Gillet, 1910 •
Copris curlettii Moretto, 2011 •
Copris davidis (Deyrolle, 1878) •
Copris davisi Nguyen-phung & Cambefort, 1986 •
Copris davisoni Waterhouse, 1891 •
Copris decellei Cambefort, 1992 •
Copris delicatus Arrow, 1931 •
Copris denticulatus Nguyen-phung, 1988 •
Copris diversus Waterhouse, 1891 •
Copris draco Arrow, 1906 •
Copris dracunculus Fereira, 1959 •
Copris druidum Heer, 1862 •
Copris dudleyi Cambefort, 1992 •
Copris eburneus Cambefort, 1992 •
Copris elpheneroides Felsche, 1910 •
Copris elphenor Klug, 1855 •
Copris erratus Lansberge, 1886 •
Copris evanidus Klug, 1855 •
Copris excisus Waterhouse, 1891 •
Copris fallaciosus Gillet, 1907 •
Copris fallax Felsche, 1910 •
Copris felschei Reitter, 1892 •
Copris fidius (Olivier, 1789) •
Copris frankenbergerianus Balthasar, 1958 •
Copris freyi Ferreira, 1961 •
Copris fricator (Fabricius, 1787) •
Copris fukiensis Balthasar, 1952 •
Copris furciceps Felsche, 1910 •
Copris garambae Cambefort, 1992 •
Copris gazellarum Gillet, 1918 •
Copris gibbulus Lansberge, 1886 •
Copris gilleti Kolbe, 1907 •
Copris gladiator Arrow, 1933 •
Copris gopheri Hubbard, 1894 •
Copris gracilis Waterhouse, 1891 •
Copris graueri Kolbe, 1914 •
Copris halffteri Matthews, 1959 •
Copris harrisi Waterhouse, 1891 •
Copris hartli Frey, 1975 •
Copris hatayamai Ochi & Araya, 1992 •
Copris hispanus (
Linnaeus, 1764) •
Copris howdeni Matthews & Halffter, 1959 •
Copris humilis Gillet, 1908 •
Copris hybridus Gillet, 1910 •
Copris igualensis Warner, 1990 •
Copris illotus Balthasar, 1942 •
Copris imitans Felsche, 1910 •
Copris inaequabilis Zhang, 1997 •
Copris incertus Say, 1835 •
Copris inemarginatus Blatchley, 1918 •
Copris inhalatus Quedenfeldt, 1884 •
Copris insidiosus Péringuey, 1901 •
Copris integer Reiche, 1847 •
Copris interioris Kolbe, 1897 •
Copris iris Sharp, 1875 •
Copris jacchoides Nguyen-phung & Cambefort, 1987 •
Copris jacchus (Fabricius, 1775) •
Copris jahi Nguyen-phung & Cambefort, 1986 •
Copris johkii Ochi & Kon, 2014 •
Copris juanmorai Mora-Aguilar & Delgado, 2015 •
Copris jucundus Gillet, 1932 •
Copris kachinensis Ochi, Kon & Kawahara, 2008 •
Copris kartlinus Kabakov, 1988 •
Copris kasagii Ochi & Kon, 1996 •
Copris kasanka Josso & Prévost, 2016 •
Copris katangae Cambefort & Nguyen-phung, 1996 •
Copris keralensis Gill, 1986 •
Copris kiuchii Masumoto, 1989 •
Copris klapperichi Balthasar, 1942 •
Copris klugi Harold, 1869 •
Copris kusakabei Ochi, Kon & Kawahara, 2005 •
Copris laeviceps Harold, 1862 •
Copris laevigatus Gillet, 1927 •
Copris laiiformis Nguyen-phung & Cambefort, 1987 •
Copris laioides Boucomont, 1932 •
Copris laius Harold, 1868 •
Copris lannathai Hanboonsong & Masumoto & Ochi, 2003 •
Copris leakeyorum Paulian, 1976 •
Copris lecontei Matthews, 1962 •
Copris leruei Josso & Prévost, 2017 •
Copris lugubris Boheman, 1858 • '''' (
Linnaeus,
1758) •
Copris macacus Lansberge, 1886 •
Copris macclevei Warner, 1990 •
Copris macer Péringuey, 1901 •
Copris maesi Ratcliffe, 1998 •
Copris magicus Harold, 1881 •
Copris manni Ochi, Kon & Bai, 2009 •
Copris marcus Gillet, 1921 •
Copris martinae Nguyen-phung, 1988 •
Copris martinezi Matthews and Halffter, 1968 •
Copris matthewsi Delgado & Kohlmann, 2001 •
Copris medogensis Zhang, 1988 •
Copris megaceratoides Waterhouse, 1891 •
Copris megasoma Matthews & Halffter, 1959 •
Copris mesacanthus Harold, 1878 •
Copris mexicanus Matthews & Halffter, 1959 •
Copris minutus (Drury, 1773) •
Copris misellus Péringuey, 1901 •
Copris moechus LeConte, 1854 •
Copris moffartsi Gillet, 1907 •
Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003 •
Copris montivagus Gillet, 1908 •
Copris morphaeus Gillet, 1932 •
Copris mourgliai Zelenka, 1993 •
Copris neglectus Moxey, 1963 •
Copris neotridens Génier & Prévost, 2017 •
Copris nepos Gillet, 1908 •
Copris nevinsoni Waterhouse, 1891 •
Copris novaki Zelenka, 1992 •
Copris nubilosus Kohlmann & Cano & Delgado, 2003 •
Copris numa Lansberge, 1886 •
Copris obenbergeri Balthasar, 1933 •
Copris obesus Boheman, 1857 •
Copris ochus (Motschulsky, 1860) •
Copris orion Klug, 1835 •
Copris orphanus Guérin-Méneville, 1847 •
Copris parapecuarius Ochi, Kon & Kawahara, 2008 •
Copris pauliani Nguyen-phung & Cambefort, 1986 •
Copris pecuarius Lewis, 1884 •
Copris pedarioides Lansberge, 1886 •
Copris phungae Cambefort, 1992 •
Copris phylax Gillet, 1908 •
Copris poggii Ochi & Kon, 2005 •
Copris potanini Semenov, 1891 •
Copris prevosti Josso, 2011 •
Copris pristinus Pierce, 1946 •
Copris pseudobootes Ferreira, 1962 •
Copris pseudochus Ochi & Kon, 2004 •
Copris pseudomoffartsi Moretto, 2011 •
Copris pseudosinicus Balthasar, 1958 •
Copris pueli Mollandin De Boissy, 1905 •
Copris punctatus Gillet, 1910 •
Copris puncticollis Boheman, 1857 •
Copris punctipennis Boucomont, 1914 •
Copris punctulatus Wiedemann, 1823 •
Copris punjabensis Gillet, 1921 •
Copris quasilaevigatus Ochi, Kon & Bai, 2007 •
Copris ramosiceps Gillet, 1921 •
Copris rebouchei Harold, 1869 •
Copris remotus Leconte, 1866 •
Copris renwarti Nguyen-phung & Cambefort, 1986 •
Copris repertus Walker, 1858 •
Copris ribbei Lansberge, 1886 •
Copris riekoae Ochi & Kon, 2003 •
Copris ritsemae Harold, 1875 •
Copris rugosus Gillet, 1908 •
Copris ruricola Balthasar, 1933 •
Copris sabinus Gillet, 1910 •
Copris sacontala Redtenbacher, 1848 •
Copris sallei Harold, 1869 •
Copris sarpedon Harold, 1868 •
Copris saundersi Harold, 1869 •
Copris schoolmeestersi Ochi & Kon, 2010 •
Copris scorpio Zelenka, 1993 •
Copris serius Nguyen-phung, 1987 •
Copris sexdentatus Thunberg, 1818 •
Copris siamensis Gillet, 1910 •
Copris siangensis Biswas, 1980 •
Copris sierrensis Matthews, 1961 •
Copris signatus Walker, 1858 •
Copris simonettai Cambefort, 1992 •
Copris simulator Nguyen-phung & Cambefort, 1986 •
Copris singularis Nguyen-phung & Cambefort, 1986 •
Copris sinicus Hope, 1842 •
Copris sinon (Olivier, 1789) •
Copris sodalis Walker, 1858 •
Copris sonensis Bui, Dumack & Bonkowski, 2018 •
Copris sorex Balthasar, 1942 •
Copris sphaeropterus Harold, 1877 •
Copris spinator Harold, 1881 •
Copris subdolus Balthasar, 1958 •
Copris subpunctatus Gillet, 1910 •
Copris subsidens Péringuey, 1901 •
Copris subterraneus Heer, 1862 •
Copris sumatrensis Gillet, 1921 •
Copris surdus Arrow, 1931 •
Copris szechouanicus Balthasar, 1958 •
Copris tetraodon Gillet, 1910 •
Copris tridentatus Solis & Kohlmann, 2003 •
Copris tripartitus Waterhouse, 1875 •
Copris truncatus Felsche, 1901 •
Copris tsukamotoi Ochi, Kon & Kawahara, 2008 •
Copris typhoeus Gerstaecker, 1884 •
Copris uenoi Hanboonsong & Masumoto & Ochi, 2003 •
Copris umbilicatus Abeille De Perrin, 1901 •
Copris urus Boheman, 1857 •
Copris usambaricus Gillet, 1908 •
Copris vankhaii Nguyen-phung, 1988 •
Copris victorini Boheman, 1857 •
Copris vietnamicus Kabakov, 1994 •
Copris vilhenai Ferreira, 1962 •
Copris vrydaghi Ferreira, 1962 •
Copris warneri Mccleve & Kohlmann, 2005 •
Copris wiesei Kolbe, 1914 •
Copris yangi Ochi, Kon & Bai, 2007 •
Copris youngai Balthasar, 1967 •
Copris zhangi Ochi, Kon & Bai, 2009 ==References==