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Rhynchotherium

Rhynchotherium is an extinct genus of proboscidea endemic to North America and Central America during the Miocene through Pliocene from 13.650 to 3.6 Ma, living for approximately 10 million years.

Taxonomy
Rhynchotherium was first described in 1868 on the basis of a lower jaw from the Miocene of Tlaxcala, Mexico. Later, the type species epithet R. tlascalae was erected for the jaw by Henry Fairfield Osborn in 1918. In 1921, a gomphothere skull from the Mt. Eden area of southern California was described as a subspecies of Trilophodon shepardi (a now-defunct combination for Mastodon shepardi), T. s. edensis, but was subsequently reassigned to Rhynchotherium. Other species subsequently assigned to Rhynchotherium included R. falconeri, R. paredensis, R. browni, and R. simpsoni. It was the closest relative to Cuvieronius, and may be ancestral to it. Lucas and Morgan (2008) reviewed the taxonomy of Rhynchotherium and concluded that only R. edensis, R. falconeri, R. paredensis, R. browni, and R. simpsoni could be confidently referred to Rhynchotherium. which it did. Misassigned speciesMastodon shepardi Leidy, 1871 • Mastodon euhypodon Cope, 1884 - likely a species of GomphotheriumTetrabelodon brevidens Cope, 1889 • Dibelodon praecursor Cope, 1893 • Rhynchotherium rectidens Osborn, 1923 • Aybelodon hondurensis Frick, 1933 • Blickotherium blicki Frick, 1933 • Rhynchotherium anguirivale Osborn, 1936 Phylogenetic position according to Mothé et al. (2016) == Life history ==
Life history
Rhynchotherium appears to have travelled in large mixed-age herds. At least two Rhynchotherium death assemblages, dubbed "elephant graveyards", are known from opposite sides of the continent, in California and Florida. The Florida site contains 3,000 individual fossils representing 38 individuals, including at least one complete adult and seven complete juveniles. The site appears to represent an area where Rhynchotherium herds repeatedly became trapped and died, potentially around the curve of a river that they periodically crossed. Some individuals may have also been washed into the site from upstream. == Palaeoecology ==
Palaeoecology
Although traditionally thought to be a browser, analysis of enamel δ13C of Rhynchotherium teeth from the Bone Valley deposits of Florida dating to the Early Pliocene suggests that they fed almost exclusively on C4 grasses. ==References==
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